Poleward Movement Research Articles

Variations in cutoff latitude during the January 2012 solar proton event and implication for the distribution of particle energy deposition

AbstractProtons in the energy range 1–20 MeV deposit most of their energy in the middle atmosphere (60–100 km). Knowledge of their magnetic latitudinal and local time distribution is crucial for determining their effect on the chemistry and dynamics in the atmosphere. Using POES 16–19 and METOP02 satellites, we investigate the latitudinal cutoff boundaries and the energy deposition during the January 2012 solar proton event. The dayside cutoff latitudes show high correlation with the Dst index even when Dst turns positive, leading to an abrupt poleward movement of more than 5°. In the same time interval, the nightside cutoff latitudes move equatorward resulting in vastly asymmetric energy deposition into the atmosphere on the dayside and nightside. The differences are sustained for almost a day in the middle atmosphere at 65° corrected geomagnetic latitude. These features cannot be taken into account by applying the frequently used GOES particle data.

Effects of Convective Gravity Wave Drag in the Southern Hemisphere Winter Stratosphere

AbstractThe excessively strong polar jet and cold pole in the Southern Hemisphere winter stratosphere are systematic biases in most global climate models and are related to underestimated wave drag in the winter extratropical stratosphere—namely, missing gravity wave drag (GWD). Cumulus convection is strong in the winter extratropics in association with storm-track regions; thus, convective GWD could be one of the missing GWDs in models that do not adopt source-based nonorographic GWD parameterizations. In this study, the authors use the Whole Atmosphere Community Climate Model (WACCM) and show that the zonal-mean wind and temperature biases in the Southern Hemisphere winter stratosphere of the model are significantly alleviated by including convective GWD (GWDC) parameterizations. The reduction in the wind biases is due to enhanced wave drag in the winter extratropical stratosphere, which is caused directly by the additional GWDC and indirectly by the increased existing nonorographic GWD and resolved wave drag in response to the GWDC. The cold temperature biases are alleviated by increased downwelling in the winter polar stratosphere, which stems from an increased poleward motion due to enhanced wave drag in the winter extratropical stratosphere. A comparison between two simulations separately using the ray-based and columnar GWDC parameterizations shows that the polar night jet with a ray-based GWDC parameterization is much more realistic than that with a columnar GWDC parameterization.

IMF effect on the polar cap contraction and expansion during a period of substorms

Abstract. The polar cap boundary (PCB) location and motion in the nightside ionosphere has been studied by using measurements from the EISCAT radars and the MIRACLE magnetometers during a period of four substorms on 18 February 2004. The OMNI database has been used for observations of the solar wind and the Geotail satellite for magnetospheric measurements. In addition, the event was modelled by the GUMICS-4 MHD simulation. The simulation of the PCB location was in a rather good agreement with the experimental estimates at the EISCAT longitude. During the first three substorm expansion phases, neither the local observations nor the global simulation showed any poleward motions of the PCB, even though the electrojets intensified. Rapid poleward motions of the PCB took place only in the early recovery phases of the substorms. Hence, in these cases the nightside reconnection rate was locally higher in the recovery phase than in the expansion phase. In addition, we suggest that the IMF Bz component correlated with the nightside tail inclination angle and the PCB location with about a 17-min delay from the bow shock. By taking the delay into account, the IMF northward turnings were associated with dipolarizations of the magnetotail and poleward motions of the PCB in the recovery phase. The mechanism behind this effect should be studied further.

Some poleward movement of British native vascular plants is occurring, but the fingerprint of climate change is not evident

Recent upperward migration of plants and animals along altitudinal gradients and poleward movement of animal range boundaries have been confirmed by many studies. This phenomenon is considered to be part of the fingerprint of recent climate change on the biosphere. Here I examine whether poleward movement is occurring in the vascular plants of Great Britain. The ranges of plants were determined from detection/non-detection data in two periods, 1978 to 1994 and 1995 to 2011. From these, the centre of mass of the population was calculated and the magnitude and direction of range shifts were determined from movements of the centre of mass. A small, but significant, northward movement could be detected in plants with expanding ranges, but not among declining species. Species from warmer ranges were not more likely to be moving northward, nor was dispersal syndrome a predictor of migration success. It is concluded that simply looking at northward movement of species is not an effective way to identify the effect of climate change on plant migration and that other anthropogenic changes obscure the effect of climate.

Boreal summer intraseasonal variability simulated in the NCEP climate forecast system: insights from moist static energy budget and sensitivity to convective moistening

The NCEP Climate Forecast System (CFS) with the relaxed Arakawa Schubert (RAS, hereafter referred to as CTRL) convection scheme of Moorthi and Suarez exhibits better performance in representing boreal summer tropical intraseasonal variability as compared with a simulation using simplified Arakawa–Schubert scheme. The intraseasonal moist static energy (MSE) budget is analyzed in this version of the CFS model (CTRL), which produces realistic eastward and northward propagation characteristics. The moist and thermodynamic processes involved in the maintenance and propagation of the poleward moving intraseasonal oscillation (ISO) disturbances are examined here. Budget diagnostics show that horizontal MSE advection is the principal component of the budget, contributing to the poleward movement of the convection. The injection of MSE moistens the atmosphere north of the convective area causing the poleward movement of convection by destabilization of the atmosphere. The moistening process is mainly contributed by the climatological wind acting on the anomalous moisture gradient as confirmed from the examination of moisture advection equation. While surface enthalpy fluxes (consisting of radiative and surface turbulent heat fluxes) maintain the ISO anomalies, they oppose the MSE tendency due to horizontal advection thus regulating the poleward propagation characteristics. In addition, the model results show that wind–evaporation feedback dominates over cloud–radiation feedback for ISO propagation; this is in contrast to our estimates using the newly available European Centre for Medium Range Weather Forecasts Interim reanalysis. Sensitivity experiments suggest that intraseasonal variability in the CFS model with the RAS scheme is highly sensitive to the parameterization of both the shallow convection and the convective rain evaporation and downdrafts. Removal of these components adversely affects the propagation characteristics and greatly reduces the amplitude of intraseasonal variability. Our results support the primary importance of the moisture preconditioning ahead of the ISO and the physical relationship between moisture and precipitation. For realistic ISO simulations, models need to represent these features appropriately.

Long-term responses of North Atlantic calcifying plankton to climate change

Climate change is threatening marine biodiversity in two ways—temperature increases and acidification. This study demonstrates that from 1960 to 2009 North Atlantic calcifying plankton primarily responded to temperature changes. Plankton communities showed an abrupt shift circa 1996, a time of a substantial temperature increase, and some taxa exhibited a poleward movement in agreement with expected biogeographical changes under ocean warming. Although acidification may become a serious threat to marine calcifiers, over the study period the primary driver of spatial distribution was ocean temperature.

The Breeding Ranges of Central European and Arctic Bird Species Move Poleward

BackgroundClimatic warming predicts that species move their entire distribution poleward. Poleward movement of the ‘cold’ side of the distribution of species is empirically supported, but evidence of poleward movement at the ‘warm’ distributional side is relatively scarce.Methodology/Principal FindingFinland has, as the first country in the world, completed three national atlas surveys of breeding birds, which we here use to calculate the sizes and weighted mean latitudes of the national range of 114 southern and 34 northern bird species during three periods (1974–1979; 1986–1989; 2006–2010), each denoting species presence in approximately 3 800 10×10 km2 squares. We find strong evidence that southern species (breeding predominantly in central Europe) showed a latitudinal shift of 1.1–1.3 km/year poleward during all three pairwise comparisons between these atlases (covering 11, 20.5 and 31.5 years respectively). We find evidence of a latitudinal shift of 0.7–0.8 km/year poleward of northern boreal and Arctic species, but this shift was not found in all study periods and may have been influenced by increased effort put into the more recent surveys. Species showed no significant correlation in changes in range size and weighted mean latitude between the first (11 year) and second (20.5 year) period covered by consecutive atlases, suggesting weak phylogenetic signal and little scope of species characteristics in explaining latitudinal avian range changes.ConclusionsExtinction-driven avian range changes (at the ‘warm’ side) of a species' distribution occur at approximately half the rate of colonisation-driven range changes (at the ‘cold’ side), and its quantification therefore requires long-term monitoring data, possibly explaining why evidence for such changes is currently rare. A clear latitudinal shift in an assemblage of species may still harbour considerable temporal inconsistency in latitudinal movement on the species level. Understanding this inconsistency is important for predictive modelling of species composition in a changing world.

Measurements of the movement of the jet streams at mid-latitudes, in the Northern and Southern Hemispheres, 1979 to 2010

Abstract. Previous studies have shown that the mean latitude of the sub-tropical jet streams in both hemispheres have shifted toward the poles over the last few decades. This paper presents a study of the movement of both the subtropical and Polar fronts, the location of the respective jet streams, between 1979 and 2010 at mid-latitudes, using total ozone measurements to identify the sharp horizontal boundary that occurs at the position of the fronts. Previous studies have shown that the two fronts are the boundaries of three distinct regimes in the stratosphere, corresponding to the Hadley, Ferrel, and polar meridionally overturning circulation cells in the troposphere. Over the period of study the horizontal area of the Hadley cell has increased at latitudes between 20 and 60 degrees while the area of the Polar cell has decreased. A linear regression analysis was performed to identify the major factors associated with the movement of the subtropical jet streams. These were: (1) changes in the Tropical land plus ocean temperature, (2) direct radiative forcing from greenhouse gases in the troposphere, (3) changes in the temperature of the lower tropical stratosphere, (4) the Quasi-Biennial Oscillation, and (5) volcanic eruptions. The dominant mechanism was the direct radiative forcing from greenhouse gases. Between 1979 and 2010 the poleward movement of the subtropical jet streams was 3.7 ± 0.3 degrees in the Northern Hemisphere and 6.5 ± 0.2 degrees in the Southern Hemisphere. Previous studies have shown that weather systems tend to follow the jet streams. The observed poleward movement in both hemispheres over the past thirty years represents a significant change in the position of the sub-tropical jet streams, which should lead to significant latitudinal shifts in the global weather patterns and the hydrologic cycle.

The observed signature of mesoscale eddies in sea surface temperature and the associated heat transport

An estimate of the signature of mesoscale motions in sea surface temperature (SST) is provided for the North Atlantic and the Southern Ocean by analysing the relationship between satellite microwave SST and multi-altimeter sea surface height (SSH) observations. After a preliminary analysis at fixed location, the study focuses on the SST/SSH relationship following eddy tracks.At fixed location, the clearest signature of mesoscale motions in SST is found in regions of large SSH variability, roughly coincident with the major fronts of the Gulf Stream and the Antarctic Circumpolar Current (ACC). Large-scale, likely atmospherically forced, variability masks a major part of the mesoscale signature in SST in more quiescent parts of the World Ocean.Following eddy tracks allows us to detect the signature of mesoscale motions in SST even in regions of weak SSH variability. The track-following analysis reveals robust westward phase shifts of the eddies' SST anomalies with respect to their rotating cores in all regions. In energetic regions, the observed intense warm-top anticyclones and cold-top cyclones are only nearly in-phase, whereas in quiet regions, weaker SST signatures are almost in quadrature with the eddies' SSH.Propagating eddies are found to flux heat poleward in the mixed-layer over a broad range of oceanic regimes and the size of this heat transport is particularly significant in the ACC region (≥0.2PW). Although eddy shedding from major currents is appealing as a mechanism for heat transport (“drift” heat transport), we find that the poleward (equatorward) motion of warm anticyclones (cold cyclones) produces a much weaker poleward heat transport in the mixed layer than that resulting from the westward phase shift between SST and SSH fluctuations (“swirl” heat transport). Associated diffusivities, modest in quiescent interiors (κ<1000m2s−1), intermediate in the ACC (1200m2s−1) and large in the Gulf Stream (≃3000m2s−1), thus primarily reflect the eddy swirl heat transport.

Genesis of twin tropical cyclones as revealed by a global mesoscale model: The role of mixed Rossby gravity waves

In this study, it is proposed that twin tropical cyclones (TCs), Kesiny and 01A, in May 2002 formed in association with the scale interactions of three gyres that appeared as a convectively coupled mixed Rossby gravity (ccMRG) wave during an active phase of the Madden‐Julian Oscillation (MJO). This is shown by analyzing observational data, including NCEP reanalysis data and METEOSAT 7 IR satellite imagery, and performing numerical simulations using a global mesoscale model. A 10‐day control run is initialized at 0000 UTC 1 May 2002 with grid‐scale condensation but no sub‐grid cumulus parameterizations. The ccMRG wave was identified as encompassing two developing and one non‐developing gyres, the first two of which intensified and evolved into the twin TCs. The control run is able to reproduce the evolution of the ccMRG wave and thus the formation of the twin TCs about two and five days in advance as well as their subsequent intensity evolution and movement within an 8–10 day period. Five additional 10‐day sensitivity experiments with different model configurations are conducted to help understand the interaction of the three gyres, leading to the formation of the TCs. These experiments suggest the improved lead time in the control run may be attributed to the realistic simulation of the ccMRG wave with the following processes: (1) wave deepening (intensification) associated with a reduction in wavelength and/or the intensification of individual gyres, (2) poleward movement of gyres that may be associated with boundary layer processes, (3) realistic simulation of moist processes at regional scales in association with each of the gyres, and (4) the vertical phasing of low‐ and mid‐level cyclonic circulations associated with a specific gyre.

Separase Sensor Reveals Dual Roles for Separase Coordinating Cohesin Cleavage and Cdk1 Inhibition

Complete dissociation of sister chromatid cohesion and subsequent induction of poleward movement of disjoined sisters are two essential events underlying chromosome segregation; however, how cells coordinate these two processes is not well understood. Here, we developed a fluorescence-based sensor for the protease separase that mediates cohesin cleavage. We found that separase undergoes an abrupt activation shortly before anaphase onset in the vicinity of chromosomes. This activation profile of separase depends on the abilities of two of its binding proteins, securin and cyclin B1, to inhibit its protease activity and target it to chromosomes. Subsequent to its proteolytic activation, separase then binds to and inhibits a subset of cyclin B1-cdk1, which antagonizes cdk1-mediated phosphorylation on chromosomes and facilitates poleward movement of sisters in anaphase. Therefore, by consecutively acting as a protease and a cdk1 inhibitor, separase coordinates two key processes to achieve simultaneous and abrupt separation of sister chromatids.

COPI-mediated membrane trafficking is required for cytokinesis inDrosophilamale meiotic divisions

The coatomer protein complex, COPI, mediates retrograde vesicle transport from the Golgi apparatus to the ER. Here, we investigated the meiotic phenotype of Drosophila melanogaster spermatocytes expressing dsRNA of 52 genes encoding membrane-trafficking-related factors. We identified COPI as an essential factor for male meiosis. In Drosophila male meiotic divisions, COPI is localized in the ER-Golgi intermediate compartment of tER-Golgi units scattered throughout the spermatocyte cytoplasm. Prior to chromosome segregation, the vesicles assemble at the spindle pole periphery through a poleward movement, mediated by minus-end motor dynein along astral microtubules. At the end of each meiotic division, COPI-containing vesicles are equally partitioned between two daughter cells. Our present data strongly suggest that spermatocytes possess a regulatory mechanism for equal inheritance of several types of membrane vesicles. Using testis-specific knockdown of COPI subunits or the small GTPase Arf or mutations of the γCOP gene, we examined the role of COPI in male meiosis. COPI depletion resulted in the failure of cytokinesis, through disrupted accumulation of essential proteins and lipid components at the cleavage furrow region. Furthermore, it caused a reduction in the number of overlapping central spindle microtubules, which are essential for cytokinesis. Drosophila spermatocytes construct ER-based intracellular structures associated with astral and spindle microtubules. COPI depletion resulted in severe disruption of these ER-based structures. Thus, we propose that COPI plays an important role in Drosophila male meiosis, not only through vesicle transport to the cleavage furrow region, but also through the formation of ER-based structures.

Identification of a TPX2-Like Microtubule-Associated Protein in Drosophila

Chromosome segregation during mitosis and meiosis relies on the spindle and the functions of numerous microtubule-associated proteins (MAPs). One of the best-studied spindle MAPs is the highly conserved TPX2, which has been reported to have characteristic intracellular dynamics and molecular activities, such as nuclear localisation in interphase, poleward movement in the metaphase spindle, microtubule nucleation, microtubule stabilisation, microtubule bundling, Aurora A kinase activation, kinesin-5 binding, and kinesin-12 recruitment. This protein has been shown to be essential for spindle formation in every cell type analysed so far. However, as yet, TPX2 homologues have not been found in the Drosophila genome. In this study, I found that the Drosophila protein Ssp1/Mei-38 has significant homology to TPX2. Sequence conservation was limited to the putative spindle microtubule-associated region of TPX2, and intriguingly, D-TPX2 (Ssp1/Mei-38) lacks Aurora A- and kinesin-5-binding domains, which are highly conserved in other animal and plant species, including many insects such as ants and bees. D-TPX2 uniformly localised to kinetochore microtubule-enriched regions of the metaphase spindle in the S2 cell line, and it had microtubule binding and bundling activities in vitro. In comparison with other systems, the contribution of D-TPX2 to cell division seems to be minor; live cell imaging of microtubules and chromosomes after RNAi knockdown identified significant delay in chromosome congression in only 18% of the cells. Thus, while this conserved spindle protein is present in Drosophila, other mechanisms may largely compensate for its spindle assembly and chromosome segregation functions.

Wind Speed Changes of North Atlantic Tropical Cyclones Preceding Landfall

AbstractLandfalling tropical cyclones have been extensively researched, especially their degradation upon coming ashore and the hazardous weather they create along coastlines and farther inland. Many of the factors that weaken storms over land could begin to act when they are still at sea, yet prelandfall intensity change remains poorly documented. This study examined the consistency of prelandfall changes in tropical cyclone intensity throughout the North Atlantic Ocean from 1950 to present. On average, cyclones intensified by 3 m s−1 less in their final 12 h before landfall than in the 12 h preceding those, on both mainland shores and in the Greater Antilles, while smaller islands showed no such change. Contributions of increasing storm age and poleward movement to this reduction were negligible. Two possible contributing factors to declining intensification before landfall were examined. The presence of a spring–neap tidal pattern of ~2 m s−1 in the prelandfall decline is consistent with tidal mixing influencing ocean temperature on the continental shelf. The tendency for the decline to be greater at night suggested the possible influence of solar heating on tropical cyclones near shore.

Air temperature and winter mortality: Implications for the persistence of the invasive mussel, Perna viridis in the intertidal zone of the south-eastern United States

Global climate change and invasive species represent two of the biggest threats to the environment. Biological communities are responding to global climate change through poleward shifts in distribution, and changes in abundance and phenology of both native and non-native species. An increase in the frequency and magnitude of extreme weather events is predicted with global climate change. Much is known about mortality events of marine organisms in relation to warm thermal stress with relatively little known about cold thermal stress, particularly in the tropics. Intertidal species are particularly susceptible to fluctuations in aerial conditions and many are considered indicators of climate change. Perna viridis is a recent invader to the United States where it fouls hard substrates and soft sediment habitats. During winter 2007/08, a mortality event was observed for P. viridis across Tampa Bay, Florida. This mortality event coincided with extreme weather conditions when air temperatures dropped below 2 °C for a period of 6 h during low water. The minimum air temperature recorded was 0.53 °C. During this period water temperature remained relatively constant (~ 20 °C). We provide strong evidence supporting the hypothesis that thermal stress relating to exposure to cold air temperatures during emersion was the primary factor underpinning the mortality event. Similar mortality events occurred in 2009 and 2010, also coinciding with prolonged exposure to low air temperatures. In the short term, weather may be responsible for the temporary trimming back of populations at the edge of their geographic range but in the longer-term, it is expected that climate warming will trigger the poleward movement of both native and non-native species potentially facilitating biotic homogenisation of marine communities. The challenge now is to devise adaptive management strategies in order to mitigate any potential negative impacts to native biodiversity.

The influence of climate variability and change on the ecosystems of the Barents Sea and adjacent waters: Review and synthesis of recent studies from the NESSAS Project

The Norwegian Component of the Ecosystem Studies of Sub-Arctic Seas (NESSAS) was funded by the Research Council of Norway from 2005 to 2008. Its aim was to quantify the impact of climate variability on the structure and function of the marine ecosystem of the Barents Sea and adjacent waters in order to predict the ecosystem responses to possible future climate change and their possible economic impact. This paper reviews research highlights dealing with climate forcing and its influence. New insights were provided on the role of large-scale atmospheric forcing on the physical oceanography including the effect of Arctic and Atlantic cyclones on the variability of the ice extent in the Barents Sea and the non-linear response of the sub-polar gyre to North Atlantic Oscillation (NAO) forcing. In addition, the NAO was also shown to influence the biology, for example shrimp recruitment in the Barents Sea and primary production in the Nordic Seas, with the strength and sign of the correlations being spatially dependent. The importance of longer term climate variability in the form of the Atlantic Multidecadal Oscillation (60–80 years period) was stressed, as it leads to significant changes in fish production, shifts in distribution and changes in spawning sites in the Barents Sea as well as other northern Atlantic ecosystems. Results from comparative studies between several US ecosystems and those of the Norwegian and Barents Seas are presented, including evidence of increased primary productivity in the Barents Sea in recent years and the poleward movement of zooplankton and fish. In addition recruitment patterns of major demersal and pelagic species in the Barents Sea generally show synchrony with those in the Gulf of Maine indicating a common external forcing. Possible ecosystem scenarios for the Barents Sea under anthropogenic-induced future climate change were developed including expectations of structural and functional changes due to distributional changes of many species. Of particular note is the likelihood of increases in the productivity of several fish species, including cod and herring, which potentially could result in higher fisheries yields.

Comment on “The importance of auroral features in the search for substorm onset process” by Syun-Ichi Akasofu, A. T. Y. Lui, and C.-I. Meng

Comment on “The importance of auroral features in the search for substorm onset process” by Syun-Ichi Akasofu, A. T. Y. Lui, and C.-I. Meng

Cytoplasmic dynein is responsible for the rapid poleward motions seen during mitosis in animals

Cytoplasmic dynein is responsible for the rapid poleward motions seen during mitosis in animals

The Dam1 ring binds to the E-hook of tubulin and diffuses along the microtubule

There has been much effort in recent years aimed at understanding the molecular mechanism by which the Dam1 kinetochore complex is able to couple microtubule depolymerization to poleward movement. Both a biased diffusion and a forced walk model have been proposed, and several key functional aspects of Dam1-microtubule binding are disputed. Here, we investigate the elements involved in tubulin-Dam1 complex interactions and directly visualize Dam1 rings on microtubules in order to infer their dynamic behavior on the microtubule lattice and its likely relevance at the kinetochore. We find that the Dam1 complex has a preference for native tubulin over tubulin that is lacking its acidic C-terminal tail. Statistical mechanical analysis of images of Dam1 rings on microtubules, applied to both the distance between rings and the tilt angle of the rings with respect to the microtubule axis, supports a diffusive ring model. We also present a cryo-EM reconstruction of the Dam1 ring, likely the relevant assembly form of the complex for energy coupling during microtubule depolymerization in budding yeast. The present studies constitute a significant step forward by linking structural and biochemical observations toward a comprehensive understanding of the Dam1 complex.

Long‐term shifts in abundance and distribution of a temperate fish fauna: a response to climate change and fishing practices

ABSTRACTAim South‐eastern Australia is a climate change hotspot with well‐documented recent changes in its physical marine environment. The impact on and temporal responses of the biota to change are less well understood, but appear to be due to influences of climate, as well as the non‐climate related past and continuing human impacts. We attempt to resolve the agents of change by examining major temporal and distributional shifts in the fish fauna and making a tentative attribution of causal factors.Location Temperate seas of south‐eastern Australia.Methods Mixed data sources synthesized from published accounts, scientific surveys, spearfishing and angling competitions, commercial catches and underwater photographic records, from the ‘late 1800s’ to the ‘present’, were examined to determine shifts in coastal fish distributions.Results Forty‐five species, representing 27 families (about 30% of the inshore fish families occurring in the region), exhibited major distributional shifts thought to be climate related. These are distributed across the following categories: species previously rare or unlisted (12), with expanded ranges (23) and/or abundance increases (30), expanded populations in south‐eastern Tasmania (16) and extra‐limital vagrants (4). Another 9 species, representing 7 families, experienced longer‐term changes (since the 1800s) probably due to anthropogenic factors, such as habitat alteration and fishing pressure: species now extinct locally (3), recovering (3), threatened (2) or with remnant populations (1). One species is a temporary resident periodically recruited from New Zealand. Of fishes exhibiting an obvious poleward movement, most are reef dwellers from three Australian biogeographic categories: widespread southern, western warm temperate (Flindersian) or eastern warm temperate (Peronian) species.Main conclusions Some of the region's largest predatory reef fishes have become extinct in Tasmanian seas since the ‘late 1800s’, most likely as a result of poor fishing practices. In more recent times, there have been major changes in the distribution patterns of Tasmanian fishes that correspond to dramatic warming observed in the local marine environment.