Interactions between societies and their natural environment have led to the development of complex traditional agroecosystems with a high level of agrodiversity (Brookfield 2001) linked to landscape mosaics which favour sustainability (Perfecto and Vandermeer 2010). In the Mediterranean region, ecological and evolutionary interactions since the Neolithic between humans and biodiversity have produced coupled bio-cultural landscapes (Blondel 2006). Domestication in that region took place through selection of biological traits for a large diversity of crops including cereals, pulses and trees (Zohary and Spiegel-Roy 1975). Crop intraspecific diversity in traditional farming systems contributes to risk avoidance strategies through the selection of adapted agronomic properties. Crop diversity also relates to sociocultural values and functions such as food preferences, ceremonial uses, identity or patrimony (Gibson 2009; Heckler and Zent 2008), as well as levels of interest within each society among individuals (Emperaire and Peroni 2007) and the reproductive biology of specific crops. Mediterranean fruit trees are allogamous and vegetative propagation is a major method for selecting and maintaining advantageous traits (Zohary and Spiegel-Roy 1975). Feral or wild crop relatives of Ficus carica, Olea europaea or Castanea sativa are preserved in traditional agroecosystems for a variety of purposes ranging from religious to practical uses, favoring gene flow between the wild or feral and clonally propagated varieties. Sexual recombinations between wild, feral and clonally propagated varieties favor the emergence of new varieties (Aumeeruddy-Thomas 2010; Aumeeruddy-Thomas et al. 2012). Studies pertaining to other vegetatively propagated crops, especially tubers in tropical regions, demonstrate the complementary role of vegetative and sexual reproduction in producing new cultivars (Caillon et al. 2006; McKey et al. 2010). In the Mediterranean region, figs, olives, dates, grapes and almonds were the first perennial plant species to be domesticated (Zohary and Hopf 2000). Compared to the roles of cereals the importance of tree interspecific and intraspecific diversity for the development of Mediterranean societies has been little discussed (Bouby and Ruas 2012). The existence of 1275 olive cultivars described to date in the Mediterranean region (Bartolini et al. 1998) shows the importance of olive intraspecific diversity at the regional level. In Morocco, 194 fig and 60 olive genotypes are known (Khadari et al. 2008b; Achtak et al. 2010). Intraspecific diversity of fruit trees in the Mediterranean region has been attributed to their cultural, economic and religious importance since ancient times. Condit (1947) monograph on the fig provides a comprehensive account of the importance of Ficus carica in legends and folklore. The sociocultural importance of fruit trees is discussed by numerous authors in classical texts on agronomy (e.g., Plinius Natural History and Columnella De Re Rustica) and in historical and ethnobotanical studies (Gsell 1914; Bellakhdar 2003). Nevertheless, no comprehensive study examines the underlying sociocultural Electronic supplementary material The online version of this article (doi:10.1007/s10745-012-9471-x) contains supplementary material, which is available to authorized users.
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