Conservation Of Old-growth Forests Research Articles

Drivers of wood-inhabiting fungal diversity in European and Oriental beech forests.

The hyperdiverse wood-inhabiting fungi play a crucial role in the global carbon cycle, but often are threatened by deadwood removal, particularly in temperate forests dominated by European beech (Fagus sylvatica) and Oriental beech (Fagus orientalis). To study the impact of abiotic drivers, deadwood factors, forest management and biogeographical patterns in forests of both beech species on fungal composition and diversity, we collected 215 deadwood-drilling samples in 18 forests from France to Armenia and identified fungi by meta-barcoding. In our analyses, we distinguished the patterns driven by rare, common, and dominant species using Hill numbers. Despite a broad overlap in species, the fungal composition with focus on rare species was determined by Fagus species, deadwood type, deadwood diameter, precipitation, temperature, and management status in decreasing order. Shifting the focus on common and dominant species, only Fagus species, both climate variables and deadwood type remained. The richness of species within the deadwood objects increased significantly only with decay stage. Gamma diversity in European beech forests was higher than in Oriental beech forests. We revealed the highest gamma diversity for old-growth forests of European beech when focusing on dominant species. Our results implicate that deadwood retention efforts, focusing on dominant fungi species, critical for the decay process, should be distributed across precipitation and temperature gradients and both Fagus species. Strategies focusing on rare species should additionally focus on different diameters and on the conservation of old-growth forests.

Forest conservation through protection of old growth: the case of Nova Scotia

Old-growth forests are both rare and special ecosystems across most of the world. Policies to protect and possibly enhance and increase them are plentiful and diverse across nations and sub-national jurisdictions. Nova Scotia has had a policy on conservation of old-growth forests on public lands in the province since 1999. The second such policy dated 2012 was recently replaced by a revised policy in 2022. After presenting knowledge based on selected scientific literature about old-growth forests in Nova Scotia, the paper describes the improvements made in the 2022 policy. These include: (a) a more-nuanced set of operational definitions; (b) a commitment to protect all old-growth forest on public lands, whether currently identified or not; (c) robust replacement provisions in the rare event that the provincial government chooses to allow provincially significant infrastructure to be built on public land supporting old-growth forest; and (d) a renewed commitment to work with private landowners on their aspirations to conserve old-growth forest. The new policy, adopted in August 2022, also contains a commitment to a public review and possible renewal by August 2027.

I boschi vetusti nel Parco del Pollino: situazione attuale e prospettive future

Old-growth forests have a primary environmental and ecological value, as they closely recall primeval forests and the natural forest landscape that was spread throughout Italy and Europe before man intervention heavily altered the functioning of most natural ecosystems. These ecosystems are more and more recognized as keystones of environmental policies (i.e., the EU Biodiversity Strategy 2030) aimed to preserve biodiversity and mitigate the impacts of climate change. A recent report highlights that Europe’s old-growth forests account for only 3 percent of the overall forest surface. The majority of italian old-growth forests are located along the Apennines, especially in the Pollino, Sila and Aspromonte National Parks, in remote montane areas characterized by irregular topographical conditions (i.e., high slopes and/or shallow soils) which hamper the access and exploitation of these less productive stands, thus allowing the survival of old-growth forests for many centuries. The ongoing climate change is a significant challenge to the conservation of old-growth forests, particularly in areas such as the Mediterranean basin, where an intensification of extreme climate events is expected. Heat waves can exacerbate drought’s negative impacts on tree growth and survival, leading to reduced woody carbon stocks and biodiversity in old-growth forests. To date, there is poor understanding of such highly valuable and extremely fragile natural ecosystems in Italy. Insights are available based on data gathered as part of two research programs, funded by the Italian Ministry of the Environment and Land and Sea Protection, with the purpose of creating a Network of Old Growth Forests in Italy’s National Parks. In this study, we aimed to highlight the role and importance of old-growth forests, focusing on those located within the Pollino National Park. A comprehensive overview of the key strengths and weaknesses, that mark these peculiar forest ecosystems, was followed by a discussion of the future outlook, grounded on strategies that will ensure not only the protection and conservation of these ecosystems, but also their development and promotion.

Forest microclimate and composition mediate long-term trends of breeding bird populations.

Climate change is contributing to biodiversity redistributions and species declines. However, cooler microclimate conditions provided by old-growth forest structures compared with surrounding open or younger forests have been hypothesized to provide thermal refugia for species that are sensitive to climate warming and dampen the negative effects of warming on population trends of animals (i.e., the microclimate buffering hypothesis). In addition to thermal refugia, the compositional and structural diversity of old-growth forest vegetation itself may provide resources to species that are less available in forests with simpler structure (i.e., the insurance hypothesis). We used 8 years of breeding bird abundance data from a forested watershed, accompanied with sub-canopy temperature data, and ground- and LiDAR-based vegetation data to test these hypotheses and identify factors influencing bird population changes from 2011 to 2018. After accounting for imperfect detection, we found that for 5 of 20 bird species analyzed, abundance trends tended to be less negative or neutral at sites with cooler microclimates, which supports the microclimate buffering hypothesis. Negative effects of warming on two species were also reduced in locations with greater forest compositional diversity supporting the insurance hypothesis. We provide the first empirical evidence that complex forest structure and vegetation diversity confer microclimatic advantages to some animal populations in the face of climate change. Conservation of old-growth forests, or their characteristics in managed forests, could help slow the negative effects of climate warming on some breeding bird populations via microclimate buffering and possibly insurance effects.

Identification of Old-Growth Mediterranean Forests Using Airborne Laser Scanning and Geostatistical Analysis

The protection and conservation of old-growth forests (OGFs) are becoming a global concern due to their irreplaceability and high biodiversity. Nonetheless, there has been little research into the identification and characterization of OGFs of the oldest tree species in Mediterranean areas. We used forest inventory data, low-density airborne laser scanning (ALS) metrics, and geostatistical analysis to estimate old-growth indices (OGIs) as indicators of old-growth forest conditions. We selected a pilot area in European black pine (Pinus nigra subsp. salzmannii) ecosystems where the oldest known living trees in the Iberian Peninsula are found. A total of 756 inventory plots were established to characterize standard live tree and stand attributes. We estimated several structural attributes that discriminate old growth from younger age classes and calculated different types of OGI for each plot. The best OGI was based on mean tree diameter, standard deviation of tree diameter, and stand density of large trees (diameter > 50 cm). This index is useful for assessing old-growthness at different successional stages (young and OGFs) in Mediterranean black pine forests. Our results confirm that the estimation of OGIs based on a combination of forest inventory data, geostatistical analysis, and ALS is useful for identifying OGFs.

Aboveground forest biomass varies across continents, ecological zones and successional stages: refined IPCC default values for tropical and subtropical forests

For monitoring and reporting forest carbon stocks and fluxes, many countries in the tropics and subtropics rely on default values of forest aboveground biomass (AGB) from the Intergovernmental Panel on Climate Change (IPCC) guidelines for National Greenhouse Gas (GHG) Inventories. Default IPCC forest AGB values originated from 2006, and are relatively crude estimates of average values per continent and ecological zone. The 2006 default values were based on limited plot data available at the time, methods for their derivation were not fully clear, and no distinction between successional stages was made. As part of the 2019 Refinement to the 2006 IPCC Guidelines for GHG Inventories, we updated the default AGB values for tropical and subtropical forests based on AGB data from >25 000 plots in natural forests and a global AGB map where no plot data were available. We calculated refined AGB default values per continent, ecological zone, and successional stage, and provided a measure of uncertainty. AGB in tropical and subtropical forests varies by an order of magnitude across continents, ecological zones, and successional stage. Our refined default values generally reflect the climatic gradients in the tropics, with more AGB in wetter areas. AGB is generally higher in old-growth than in secondary forests, and higher in older secondary (regrowth >20 years old and degraded/logged forests) than in young secondary forests (⩽20 years old). While refined default values for tropical old-growth forest are largely similar to the previous 2006 default values, the new default values are 4.0–7.7-fold lower for young secondary forests. Thus, the refined values will strongly alter estimated carbon stocks and fluxes, and emphasize the critical importance of old-growth forest conservation. We provide a reproducible approach to facilitate future refinements and encourage targeted efforts to establish permanent plots in areas with data gaps.

Knowledge and conservation of Old-Growth Forests: a key issue to face global changes. The case study of Strâmbu-Băiuț - Maramureş (Eastern Carpathians, Romania)

Over the last three decades, worldwide forests have become increasingly the focus of societal and scientific interest. The critical ecosystem services they provide are considered as key elements to mitigate harmful effects of global changes. Most of recent studies highlighted the highest effectiveness of the so-called ‘primary’, ‘natural’ and ‘old-growth forests’ in this struggle due to their specific features (biodiversity and carbon storage capacity). However, in Europe, these forests which represent less than 1% of total forest cover are threatened by unsustainable human activity. Mainly based on pure ecological investigations, most of recent studies have rarely integrated human dimensions so far. Several outstanding issues remain to be addressed about the real nature of the “remaining natural European forests”. Tackling this issue is crucial to know which forest must be conserved and how. In order to improve (i) knowledge about and (ii) conservation and /or sustainable management of such forests, we carried out an integrative and innovative research combining natural, social and human science approaches on a recently classified UNESCO primary beech forest located in the Maramures county (Eastern Carpathians, Romania). First results highlighted that far from being untouched, this current high-value forest results from a long-term co-evolution with local communities. It implies a deeper understanding of the complex interaction between ecological and anthropogenic legacies is mandatory to improve and grant the preservation and sustainability of old-growth forests.

Comparative models disentangle drivers of fruit production variability of an economically and ecologically important long-lived Amazonian tree

Trees in the upper canopy contribute disproportionately to forest ecosystem productivity. The large, canopy-emergent Bertholletia excelsa also supports a multimillion-dollar commodity crop (Brazil nut), harvested almost exclusively from Amazonian forests. B. excelsa fruit production, however is extremely variable within populations and years, destabilizing local harvester livelihoods and the extractive economy. To understand this variability, data were collected in Acre, Brazil over 10 years at two sites with similar climate and forest types, but different fruit production levels, despite their proximity (~ 30 km). One site consistently produced more fruit, showed less individual- and population-level variability, and had significantly higher soil P and K levels. The strongest predictor of fruit production was crown area. Elevation and sapwood area also significantly impacted fruit production, but effects differed by site. While number of wet days and dry season vapor pressure prior to flowering were significant production predictors, no climatic variables completely captured annual observed variation. Trees on the site with higher available P and K produced nearly three times more fruits, and appeared more resilient to prolonged drought and drier atmospheric conditions. Management activities, such as targeted fertilization, may shield income-dependent harvesters from expected climate changes and production swings, ultimately contributing to conservation of old growth forests where this species thrives.

Patterns of biodiversity response along a gradient of forest use in Eastern Amazonia, Brazil.

The preservation of tropical forests is increasingly at risk, including forests located within human-modified landscapes that retain high conservation value. People modify and interact with these landscapes through a wide range of uses. However, our knowledge of how different forest uses affect biodiversity is limited. Here, we analyse the responses of different taxa to four distinct categories of forest management, namely old-growth forest, Brazil nut extraction areas, reduced impact logging areas, and eucalyptus plantations. Within six independent replicates of each category, we sampled three taxa (fruit-feeding butterflies, dung beetles, and trees) in eastern Amazonia. Forests under moderate use (Brazil nut extraction and reduced-impact logging) had similar, albeit slightly lower, diversity levels relative to old-growth forests, while communities in plantations were significantly less diverse. Only 4%, 20%, and 17%, of the sampled butterfly, dung beetle, and tree species, respectively, were restricted to old-growth forests. This study provides further empirical evidence of the importance of old-growth forest conservation in the context of human-modified landscapes. It also suggests that landscape matrices integrating forest uses at varying intensities are well positioned to reconcile biodiversity conservation with the production of goods that support local livelihoods.

Secondary disturbances of low and moderate severity drive the dynamics of eastern Canadian boreal old-growth forests

Both low- and moderate-severity secondary disturbances are drivers of eastern Canadian boreal old-growth forests dynamics. Moderate-severity disturbances reflect mainly spruce budworm outbreaks. Low-severity disturbances are produced by both spruce budworm outbreaks and random events such as windthrow. Each level of disturbance severity has a specific impact on stand dynamics, and both shape the diversity of boreal old-growth forests. A regular succession of low-severity disturbances is seen as determining the dynamics of the old-growth stage (gap dynamics); however, recent studies suggest that moderate-severity secondary disturbances also play an important role in the dynamics of eastern Canadian boreal forests. This study aims to determine if eastern Canadian boreal old-growth forests are driven by a combination of low- and moderate-severity secondary disturbances. We reconstructed the 200-year disturbance history of 20 boreal old-growth stands using dendrochronological analysis. We discriminated low- from moderate-severity disturbances based on their respective influence on mean stand growth. The secondary disturbance regime of eastern Canadian boreal old-growth forests varies highly over time, reflected by disturbance peaks in the chronological record. Most peaks occurred during spruce budworm outbreaks related to both low- and moderate-severity disturbances. Between each peak, low-severity disturbances dominate. Each level of disturbance severity has specific consequences for stand dynamics. Both low and moderate secondary disturbances are drivers of forest dynamics in eastern Canadian boreal old-growth stands and shape the structural diversity of these stands. The complexity of these dynamics should be recognized in management planning to ensure the efficiency of old-growth forest conservation policies.

Value-added forest management planning: A new perspective on old-growth forest conservation in the fire-prone boreal landscape of Canada

The maintenance of old-growth stands is important for sustaining natural forest ecosystems, but fire disturbances and commonly-used timber harvest practices exert adverse impacts on the retention of old-growth forests. Forest management planning prescribes harvest levels based on the planning policy and models, but the impact of the management strategies on the retention of old-growth forests has not been well studied. The objectives of this study were to examine: a) the impact of implementing three different harvest policies on the retention of old-growth forest and b) the impact of implementing a policy of maintaining a targeted minimum of 20% old-growth area on the harvest revenue that would be generated over a long planning horizon. To simulate the implementation of these policies, we developed three strategic timber harvest-scheduling models. The first model (Model 1) maximizes harvest timber volume; Model 2 maximizes the net present value (NPV) of the timber harvested; and Model 3 maximizes the NPV of value-added products at the primary processing mills. The value-added products we considered were lumber, chips and sawdust. The models were solved for three forest management units with different fire regimes. Solutions to models that did not include a strict constraint on old-growth forest area retention did not retain the targeted level of old-growth forest over a 150-year planning horizon. When an old-growth constraint was implemented, Model 3 produced the greatest revenue with the least variation by 5-year period over the planning horizon. The probability of finding a feasible solution to our optimization Model 3 with an old-growth forest constraint increased to 0.87–1.0 compared with 0.71–0.83 using Model 1, and 0.78–0.87 using Model 2. We conclude that the value-added policy model increases the probability of sustaining the bioeconomy while preserving forest ecosystems initiated by disturbance.

Old forest policy and regulatory frameworks in Nova Scotia and New Brunswick with a comparison to British Columbia

Old growth forests (OGF) represent ecological continuity and are rich in biodiversity. These unique landscapes are becoming increasingly rare in Canada, especially within the Acadian Forest Region that covers the Maritime Provinces. Less than 1% of the total forest cover in these provinces is OGF — this is drastically lower than the 55% represented in British Columbia’s Temperate Coastal Rainforest. The authors of this paper conducted a comprehensive comparative analysis of old forest policies and regulatory frameworks in Nova Scotia, New Brunswick and British Columbia to offer recommendations for improving the conservation and stewardship of OGF in the Maritimes. The analysis revealed that in Nova Scotia there is a strong old forest policy underpinned by a quantitative scoresheet, coordinator, and database. In New Brunswick and Nova Scotia the conservation targets are up to 8% on Crown land, but neither have a regulatory framework nor have evidence of achieving these targets. British Columbia has integrated Forest Stewardship Plans that promote the retention of OGF as legally enforceable tools for licensees in Old Growth Forest Management Areas. Such an approach could prove useful in improving the restoration and conservation of OGF in the Maritime Provinces.

Disturbance and mosquito diversity in the lowland tropical rainforest of central Panama

The Intermediate Disturbance Hypothesis (IDH) is well-known in ecology providing an explanation for the role of disturbance in the coexistence of climax and colonist species. Here, we used the IDH as a framework to describe the role of forest disturbance in shaping the mosquito community structure, and to identify the ecological processes that increase the emergence of vector-borne disease. Mosquitoes were collected in central Panama at immature stages along linear transects in colonising, mixed and climax forest habitats, representing different levels of disturbance. Species were identified taxonomically and classified into functional categories (i.e., colonist, climax, disturbance-generalist, and rare). Using the Huisman-Olff-Fresco multi-model selection approach, IDH testing was done. We did not detect a unimodal relationship between species diversity and forest disturbance expected under the IDH; instead diversity peaked in old-growth forests. Habitat complexity and constraints are two mechanisms proposed to explain this alternative postulate. Moreover, colonist mosquito species were more likely to be involved in or capable of pathogen transmission than climax species. Vector species occurrence decreased notably in undisturbed forest settings. Old-growth forest conservation in tropical rainforests is therefore a highly-recommended solution for preventing new outbreaks of arboviral and parasitic diseases in anthropic environments.

Balancing water supply and old-growth forest conservation in the lowlands of south-central Chile through adaptive co-management

Ecosystem management is a conservation strategy, but there is not a standard protocol for implementation. In theory, ecosystem management will utilize the best available science to sustain social-ecological systems in the landscape by maximizing multiple ecosystem services expected from the stakeholders’ of those systems. Llancahue is a watershed (1,270 ha) that provides fresh water to the city of Valdivia (130,000 inhabitants) and protects >700 ha of old-growth forest within a severely disturbed landscape in the lowlands of south-central Chile. The native vegetation of this landscape is the threatened Valdivian Temperate Rainforests. Management of the watershed by the Universidad Austral de Chile needs to both provide timber and work to the neighboring campesinos (small poor rural land owners) who were illegally logging the forest and improve the conservation of the old-growth forests and the quantity and quality of water provided by the watershed. This paper demonstrates how adaptive management has utilized a multi-step process to improve management of the watershed. This process has included (1) understanding stakeholders’ views towards the project, (2) developing an ecosystem management plan for the watershed that balanced multiple societal demands and ecosystem functions from the watershed, and (3) monitoring the Llancahue forest and streams to ensure activities provided desired results. This paper reports results after 4 years of implementation and provides perspectives on the ecosystem management approach.

Riparian old-growth forests provide critical nesting and foraging habitat for Blakiston's fish owl Bubo blakistoni in Russia

AbstractConservation efforts for Blakiston's fish owl Bubo blakistoni in Russia are limited, partly because habitat use by these rare owls is poorly known. We therefore studied nesting and foraging habitat characteristics of Blakiston's fish owls in Primorye, Russia. We sampled habitat at 14 nest sites, 12 nest stand sites and 13 random sites; we also sampled rivers within 14 fish owl home ranges across our 20,213 km2 study area. We found that large old trees and riparian old-growth forest were the primary characteristics of nest and foraging sites, respectively. Large trees were probably used as nest sites because they have cavities large enough to accommodate these birds. Big trees are also important because they are primary sources of large woody debris in rivers, which enhances suitable habitat for salmon, the owl's primary prey. Based on habitat characteristics, nest sites were correctly distinguished from random sites 74% (Kappa = 0.48) of the time, nest stands from random sites 56% (Kappa = 0.12) of the time, and used sites from available foraging sites 68% (Kappa = 0.36) of the time. The management and conservation of old-growth forests is essential for sustaining this species because they are central to the owls' nesting and foraging behaviour. Moreover, conservation of these forests sustains habitat for many other species.

Fire‐mediated pathways of stand development in Douglas‐fir/western hemlock forests of the Pacific Northwest, USA

Forests dominated by Douglas-fir and western hemlock in the Pacific Northwest of the United States have strongly influenced concepts and policy concerning old-growth forest conservation. Despite the attention to their old-growth characteristics, a tendency remains to view their disturbance ecology in relatively simple terms, emphasizing infrequent, stand-replacing (SR) fire and an associated linear pathway toward development of those old-growth characteristics. This study uses forest stand- and age-structure data from 124 stands in the central western Cascades of Oregon to construct a conceptual model of stand development under the mixed-severity fire regime that has operated extensively in this region. Hierarchical clustering of variables describing the age distributions of shade-intolerant and shade-tolerant species identified six groups, representing different influences of fire frequency and severity on stand development. Douglas-fir trees > 400 years old were found in 84% of stands, yet only 18% of these stands (15% overall) lack evidence of fire since the establishment of these old trees, whereas 73% of all stands show evidence of at least one non-stand-replacing (NSR) fire. Differences in fire frequency and severity have contributed to multiple development pathways and associated variation in contemporary stand structure and the successional roles of the major tree species. Shade-intolerant species form a single cohort following SR fire, or up to four cohorts per stand in response to recurring NSR fires that left living trees at densities up to 45 trees/ha. Where the surviving trees persist at densities of 60-65 trees/ha, the postfire cohort is composed only of shade-tolerant species. This study reveals that fire history and the development of old-growth forests in this region are more complex than characterized in current stand-development models, with important implications for maintaining existing old-growth forests and restoring stands subject to timber management.

Exploring symbiont management in lichens

Lichens are unique among fungal symbioses in that their mycelial structures are compact and exposed to the light as thallus structures. The myriad intersections of unique fungal species with photosynthetic partner organisms (green algae in 90% of lichens) produce a wide variety of diverse shapes and colours of the fully synthesized lichen thallus when growing in nature. This characteristic complex morphology is, however, not achieved in the fungal axenic state. Even under ideal environmental conditions, the lichen life cycle faces considerable odds: first, meiotic spores are only produced on well-established thalli and often only after achieving considerable age in a stable environment, and second, even then in vivo resynthesis requires the presence of compatible algal strains where fungal spores germinate. Many lichen species have evolved a way around the resynthesis bottleneck by producing asexual propagules for joint propagation of symbionts. These different dispersal strategies ostensibly shape the population genetic structure of lichen symbioses, but the relative contributions of vertical (joint) and horizontal (independent) symbiont transmission have long eluded lichen evolutionary biologists. In this issue of Molecular Ecology, Dal Grande et al. (2012) close in on this question with the lung lichen, Lobaria pulmonaria, a flagship species in the conservation of old growth forests. By capitalizing on available microsatellite markers for both fungal and algal symbionts, they show that while vertical transmission is the predominant mode of reproduction, horizontal transmission is demonstrable and actively shapes population genetic structure. The resulting mixed propagation system is a highly successful balance of safe recruitment of symbiotic clones and endless possibilities for fungal recombination and symbiont shuffling.

Network centrality and social movement media coverage: A two-mode network analytic approach

This article examines the relationship between structural location (namely, degree centrality) and news media coverage. Our central hypothesis is that the network centrality of social movement actors is positively associated with the prevalence of actors being cited in the print news media. This paper uses two-mode data from a communication network of environmentalists in British Columbia, and examines the relationship between their structural location and the frequency by which they are cited in newsprint media with regard to particular frames (about forest conservation, environmental protest, and related issues). We asked a sample of social movement participants about their ties to a target list of relatively high profile actors (environmental activists). We turned the resulting network matrix into a bipartite graph that examined the relationships amongst the target actors vis a vis the respondents. Next we calculated point in-degree for the target actors. For the target actors we also have data from a representative sample of 957 print news articles about forestry and conservation of old growth forests in British Columbia. We compare the effects of network centrality of the target actor versus several attributes of the target actors (gender, level of radicalism, leadership status) on the amount of media coverage that each of the target actors receives. We find that network centrality is associated with media coverage controlling for actor attributes. We discuss theoretical implications of this research. Finally, we also discuss the methodological pros and cons of using a “target name roster” to construct two-mode data on social movement activists.

Living among Frequent-fire Forests: Human History and Cultural Perspectives

Murphy, A., J. Abrams, T. Daniel, and V. Yazzie. 2007. Living among frequent-fire forests: human history and cultural perspectives. Ecology and Society 12(2): 17. https://doi.org/10.5751/ES-02167-120217

The Northwest Forest Plan: Origins, Components, Implementation Experience, and Suggestions for Change

In the 1990s the federal forests in the Pacific Northwest underwent the largest shift in management focus since their creation, from providing a sustained yield of timber to conserving biodiversity, with an emphasis on endangered species. Triggered by a legal challenge to the federal protection strategy for the Northern Spotted Owl (Strix occidentalis caurina), this shift was facilitated by a sequence of science assessments that culminated in the development of the Northwest Forest Plan. The plan, adopted in 1994, called for an extensive system of late-successional and riparian reserves along with some timber harvest on the intervening lands under a set of controls and safeguards. It has proven more successful in stopping actions harmful to conservation of old-growth forests and aquatic systems than in achieving restoration goals and economic and social goals. We make three suggestions that will allow the plan to achieve its goals: (1) recognize that the Northwest Forest Plan has evolved into an integrative conservation strategy, (2) conserve old-growth trees and forests wherever they occur and (3) manage federal forests as dynamic ecosystems.