Flowers of Cochliostema odoratissimum are trimerous with three fertile stamens, three unequal antherless staminodes, and three connate carpels. The fertile stamens are on one side of the flower and united by their filaments, forming a compound structure that curves to the flower's right as the flower opens. The thecae are longitudinally dehiscent, spirally coiled, and enveloped by petaloid extensions of the filaments of the two lateral stamens contributing to the three-staminate structure. Anther wall development is of the monocotyledonous type. Tapetal raphides are formed and appear to be widespread in Commelinaceae. Also known from Philydraceae and, perhaps, Haemodoraceae, tapetal raphides and their taxonomic distribution may be of phylogenetic utility. Microsporogenesis is successive, forming both isobilateral and decussate tetrads. Pollen is shed as single binucleate grains. Each ovary locule contains ten to twelve hemianatropous, crassinucellar, bitegmic ovules on axile placentae. The micropyle is formed by both the inner integument and one side of the outer integument. Megagametophyte development is of the Polygonum type. The mature megagametophyte consists of an egg apparatus, fusion nucleus, and three antipodals, the latter showing signs of degeneration. The salient features of the floral vasculature are the same as in the few other commelinaceous species for which complete data are available. Relative to the floral vasculature in the other species, differences in the vasculature lie primarily in the presence and origin of lateral carpel bundles and in the number of sepal and ovule traces.