IN this communication the author endeavoured to summarise the main facts already known regarding the palæontological history of the Crocodilia, with full references to the principal literature of the subject. After some preliminary remarks upon the structure and distribution of the living members of the order, the leading types of each geological period were successively considered; and the paper concluded by discussing the bearing of these facts upon the evolution of the Crocodilia, as determined by Prof. Huxley in 1875. The earliest crocodilians hitherto discovered are Belodnn, Stagonolepis, and Parasuchus, from the Upper Trias—the firstnamed being met with on the Continent, in India, and in North America; the second, solely known from the yellow sandstones of Elgin; and the third, only recorded as yet from India. The Rhætic Beds and Lower and Middle Lias do not appear to have yielded any remains of this order, but numerous examples have been found in the Upper Lias, and some in a remarkable state of preservation. At present, however, the precise systematic relationships of the Liassic forms have not been very satisfactorily deter mined, and those from British deposits are especially in need of further study: there are probably two generic types, Mystriosaurus and Pelagosaurus , and, if the latest researches are to be followed, it would seem that only two species of each are definitely known. In England, according to M. Deslongchamps, two distinct forms, Mystriosaurus chapmani and Pelagosaurus brongniarti . have been continually described under the name of Teleosaurus chapmani . With the Lower Oolites, Teleosaurus proper makes its appearance, and ranges at least as far upwards as the Kimmeridge Clay, from which Mr. J. W. Hulke has described a characteristic snout ( T. megarhinus). Steneosaurus is also a Lower Oolite form, ranging to the Oxford Clay; its British representatives are some what imperfectly known, though very complete descriptions have been published of well-preserved cranial fossils from French deposits. Metriorhynchus is another genus, from the Middle and Upper Oolites, very fully elucidated by M. Deslongchamps in France, but scarcely determined hitherto in English strata. Two forms described by Prof. Phillips under the names of Steneosaurus palpebrosus (Kimmeridgian), and Steneosaurus gracilis (Portlandian) are truly referable to Metriorhynchus , and fragments agreeing specifically with some of the French Metriorhynchs are also recorded. The Upper Oolites also yield the remains of Crocodilia with comparatively short and stout skulls, and very complete specimens have been discovered in the Kimmeridge Clay both of England and the Continent. They belong to the genera Dakosaurus and Machimosaurus , the former having also been described by Sir Richard Owen under the name of Plesiosuchus . Teleosaurians occur rarely in the Wealden and Purbeck Beds—though one or two well-preserved crania of Macrorhynchus are known in Germany—and they finally disappear in the Upper Cretaceous series, where they are represented by the scanty remains of Hyposaurus and Enaliosuchus . Broad-faced crocodilians, adapted for a moie terrestrial mode of life than the Teleosaurs, occur somewhat abundantly in the Wealden and Purbeck Beds, and are represented by Goniopholis, Nannosuchus, Oweniasuchus, Theriosuchus , and a remarkably interesting genus— Bernissartia—recently described by M. Dollo from the now classical deposit of Bernissart in Belgium. The latter, though decidedly Mesosuchian, approaches the living crocodiles and alligators much more closely than any of its congeners, both in the characters of its dermal armour and in certain parts of the skull. The earliest evidence of procœlian crocodiles hitherto made known is from the Cambridge Greensand and the nearly equivalent Gosau Beds of Vienna; Prof. H. G. Seeley has described a few vertebræ, teeth, and fragments of limb-bones, and regards these as referable to at least three specific types. The Upper Cretaceous beds of France and the United States have yielded still more satisfactory remains—including skulls—howing that the Eusuchian sub-order dates back beyond Eocene times; and some of these fossils appear almost indistinguishable from the living genus Gavialis. The early Tertiary deposits, both of England and the Continent, are remarkable as affording traces of gavials, crocodiles, and alligators (or alligatoroid genera) associated together, while the three families share no common area of the earth's surface at the present time; the Eocene types, moreover, appear to be rather less differentiated than is the case in the existing fauna. In conclusion, it may be said that the abundant acquisitions of fossil Crocodilia during the last ten years have fully confirmed the views of Prof. Huxley, laid before the Geological Society in 1875; and the Wealden and Purbeck discoveries, particularly, have brought to light facts which were then little more than probable surmises based upon very fragmentary materials.
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