Beetle-killed Lodgepole Pine Research Articles

Evaluation of Compatibility between Beetle-Killed Lodgepole Pine (Pinus Contorta var. Latifolia) Wood with Portland Cement.

The compatibility of wood from mountain pine beetle (Dendroctonus ponderosa) killed lodgepole pine (Pinus contorta var. latifolia) with Portland cement was investigated based on time-since-death as a quantitative estimator, and the presence of blue-stained sapwood, brown rot, or white rot as qualitative indicators. The exothermic behavior of cement hydration, maximum heat rate, time to reach this maximum, and total heat released within a 3.5–24 h interval were used for defining a new wood-cement compatibility index (CX). CX was developed and accounted for large discrepancies in assessing wood-cement compatibility compared to the previous methods. Using CX, no significant differences were found between fresh or beetle-killed wood with respect to the suitability for cement; except for the white rot samples which reached or exceeded the levels of incompatibility. An outstanding physicochemical behavior was also found for blue-stained sapwood and cement, producing significantly higher compatibility indices.

Effects of salvage logging on avian nest survival in beetle-killed forests

Information about the immediate and potentially prolonged impacts of salvage logging is needed to develop prescriptions that maintain forest productivity, reduce the risk of catastrophic fires, and conserve biological communities. Most salvage logging studies have investigated post-fire salvage, often clearcutting, and impacts documented in these studies may be more severe than selective logging prescriptions applied to forests affected by beetle outbreaks. We evaluated the effects of selective salvage logging on avian nest survival in beetle-killed lodgepole pine (Pinus contorta) forests on the Fremont and Winema National Forests, south-central Oregon, USA, 1997–1999. Using logistic-exposure models, we found that nest survival for weak (period nest survival: 0.64; 95% CL: 0.50, 0.75) and strong (period nest survival: 0.71; 95% CL: 0.46, 0.87) cavity nesters was significantly higher than for ground nesters (period nest survival: 0.31; 95% CL: 0.18, 0.46), while nest survival for open cup nesting birds (period nest survival: 0.57; 95% CL: 0.47, 0.66) was intermediate. We did not find significant differences in nest survival for American robin (Turdus migratorius), chipping sparrow (Spizella passerine), dark-eyed junco (Junco hyemalis), dusky flycatcher (Empidonax oberholseri), mountain chickadee (Poecile gambeli), or yellow-rumped warbler (Dendroica coronata) (these six species comprised ∼80% of the sample). Our results suggest that, unlike post-fire salvage logging, selective harvesting prescriptions after beetle outbreaks may be suitable for meeting multiple management objectives, including generating a flow of commodities and maintaining avian nest survival rates.

Green-tree retention and life after the beetle: stand structure and small mammals 30 years after salvage harvesting

We report on a retrospective investigation of the impacts of salvage harvesting of lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Wats.), killed by an outbreak of mountain pine beetle (Dendroctonus ponderosae Hopk.) in the 1970s, with variable retention of Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco). Our inference to biodiversity was coniferous stand structure and four mammal species: the southern red-backed vole (Myodes gapperi Vigors), common shrew (Sorex cinereus Kerr), red squirrel (Tamiasciurus hudsonicus Erxleben) and northern flying squirrel (Glaucomys sabrinus Shaw). We tested hypotheses that, at 30 years after salvage harvest of beetle-killed lodgepole pine trees, (1) abundance and diversity of stand structure, and (2) abundance of mammal species, will increase with higher levels of green-tree retention (GTR). Stand structure attributes and small mammals were sampled during 2005–2008 in young pine stands, with a range of GTR seed-trees (none, dispersed, and aggregated Douglas-fir), and uncut forest in south-central British Columbia, Canada. Diameters and heights of Douglas-fir and lodgepole pine and basal area of total conifers supported hypothesis (1). Mean abundance of the red-backed vole was consistently higher (2.3 to 6.4 times) in the uncut forest than other stands. Overall mean patterns of abundance for common shrews, red squirrels, and northern flying squirrels were similar among treatment stands. Mean abundance of the red-backed vole supported hypothesis (2), but numbers of the other three species did not. There is “life after the beetle” at 30 years after salvage harvesting, and this was enhanced by GTR.

The Effect of Varying Organosolv Pretreatment Chemicals on the Physicochemical Properties and Cellulolytic Hydrolysis of Mountain Pine Beetle-Killed Lodgepole Pine

Mountain pine beetle-killed lodgepole pine (Pinus contorta) chips were pretreated using the organosolv process, and their ease of subsequent enzymatic hydrolysis was assessed. The effect of varying pretreatment chemicals and solvents on the substrate's physicochemical characteristics was also investigated. The chemicals employed were MgCl2, H2SO4, SO2, and NaOH, and the solvents were ethanol and butanol. It was apparent that the different pretreatments resulted in variations in both the chemical composition of the solid and liquid fractions as well in the extent of cellulolytic hydrolysis (ranging from 21% to 82% hydrolysis after 12 h). Pretreatment under acidic conditions resulted in substrates that were readily hydrolyzed despite the apparent contradiction that pretreatment under alkaline conditions resulted in increased delignification (approximately 7% and 10% residual lignin for alkaline conditions versus 17% to 19% for acidic conditions). Acidic pretreatments also resulted in lower cellulose degree of polymerization, shorter fiber lengths, and increased substrate porosity. The substrates generated when butanol/water mixtures were used as the pretreatment solvent were also hydrolyzed more readily than those generated with ethanol/water. This was likely due to the limited miscibility of the solvents resulting in an increased concentration of pretreatment chemicals in the aqueous layer and thus a higher pretreatment severity.

Temporal change in wood quality attributes in standing dead beetle-killed lodgepole pine

The number and size of checks, wood moisture content, extent of blue-stain, rot and decay was examined by stem analysis in 360 mature standing beetle-killed lodgepole pines. Trees came from three areas (Burns Lake, Quesnel, and Vanderhoof) in Central British Columbia. Each area was represented by 14 to 16 sampling areas (stands) distributed evenly across three soil moisture regimes (dry, mesic, wet). Year of death was estimated from tree ring-analysis, local knowledge and insect and disease survey maps. An approximately equal number of trees had been dead for one or two years, three or four years, or for five or more years. During the first five years since death by beetle attack the number of checks per tree increased from 2.5 to 10.2 and the average depth of checks increased from 4.3 cm to 5.1 cm. Checks were deeper, wider, and longer on the drier sites than on mesic and wet sites. Moisture content of sapwood and heartwood was near the fibre saturation point (ca 30%) one year after death and continued to decrease at a rate of approximately 1.7% per year. Both the incidence and the extent (relative to basal area) of rot and decay increased significantly with time since death. All trees had an extensive blue-stain discoloration. Deterioration of wood quality was fastest during the first two years after a beetle attack. Key words: wood checks, moisture content, blue-stain, wood quality, spiral grain, bark beetle

Acid‐catalyzed steam pretreatment of lodgepole pine and subsequent enzymatic hydrolysis and fermentation to ethanol

Utilization of ethanol produced from biomass has the potential to offset the use of gasoline and reduce CO(2) emissions. This could reduce the effects of global warming, one of which is the current outbreak of epidemic proportions of the mountain pine beetle (MPB) in British Columbia (BC), Canada. The result of this is increasing volumes of dead lodgepole pine with increasingly limited commercial uses. Bioconversion of lodgepole pine to ethanol using SO(2)-catalyzed steam explosion was investigated. The optimum pretreatment condition for this feedstock was determined to be 200 degrees C, 5 min, and 4% SO(2) (w/w). Simultaneous saccharification and fermentation (SSF) of this material provided an overall ethanol yield of 77% of the theoretical yield from raw material based on starting glucan, mannan, and galactan, which corresponds to 244 g ethanol/kg raw material within 30 h. Three conditions representing low (L), medium (M), and high (H) severity were also applied to healthy lodgepole pine. Although the M severity conditions of 200 degrees C, 5 min, and 4% SO(2) were sufficiently robust to pretreat healthy wood, the substrate produced from beetle-killed (BK) wood provided consistently higher ethanol yields after SSF than the other substrates tested. BK lodgepole pine appears to be an excellent candidate for efficient and productive bioconversion to ethanol.

Bird Response to Beetle-Killed Lodgepole Pine

Bird Response to Beetle-Killed Lodgepole Pine