Abstract

SummaryCarlquist, S.: Wood and stem anatomy ofSarcobatus (Caryophyllales):systematic and ecological implications. – Taxon 49: 27‐34. 2000. – ISSN 0040‐0262.Quantitative and qualitative data are given forSarcobatus vermiculatus(Hook.) Torr, stem anatomy in order to provide information relevant to Behnke's segregation ofSarcobatusfromChenopodiaceaeand his idea thatSarcobataceaebelong in suborderPhytolaccineaerather than suborderChenopodiineae. Sarcobatushas highly xeromorphic wood such as in xericChenopodiaceae(presence of vasicentric tracheids, narrow vessels, short vessel elements, numerous vessels per group, helical sculpturing of vessel walls), but thesefeaturescould have evolved independently in both suborders. Nonbordered perforation plates, present inSarcobatus, are widespread inCaryophyllaless.str. and may prove to be a defining feature for the recently expandedCaryophyllalestogether with the neighbouring orderSantalales.A single‐layered lateral meristem occurs in genera ofChenopodiinaewith successive cambia, but also in genera ofPhytolaccineae.Likewise, raylessness occurs in both suborders. Stem and wood anatomy does not provide decisive evidence for placement ofSarcobatusinPhytolaccineae, but it does not provide any evidence against that treatment.

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