Abstract
The nutritional status of the soils of a series of coniferous woodland clearings in the Netherlands was found to be intermediate between that of the soil in a dune area with a climax population of Chamaenerion angustifolium and that of the edge (and undergrowth) of a Quercus robur stand. In the dune area the greater part of the NPK was present as an internal pool in the plant tissues of C. angustifolium enabling the population of this species to persist and inhibiting further succession through limiting the availability of NPK. It is suggested that succession in clearings might stagnate in the same way in the absence of new tree growth. Because C. angustifolium is capable of persisting as a population under poor N conditions, it is unsuitable as an indicator species for nitrate-rich soils. The same may be true for Senecio sylvaticus, the other character species of the association Epilobio-Senecionetum sylvatici.
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