Nomenclator of Bivalve Families with a Classification of Bivalve Families

2,604 names at the rank of subtribe, tribe, subfamily, family and superfamily have been proposed for Recent and fossil gastropods, and another 35 for monoplacophorans. All names are listed in a nomenclator giving full bibliographical reference, date of publication, typification, and their nomenclatural availability and validity under the International Code of Zoological Nomenclature. Another 790 names, established for categories above the familygroup (infraorder to subclass) are listed separately. A fully ranked, hierarchical classification summarizes recent advances in the phylogeny of the Gastropoda and Monoplacophora. In all, the classification recognizes as valid a total of 721 gastropod families, of which 245 are known exclusively as fossils and 476 occur in the Recent with or without a fossil record; and 20 monoplacophoran families, of which 1 only occurs as Recent.Nomenclatural acts in this work: Amberleya bathonica Cox & Arkell, 1950, fixed as type species of Amberleya J. Morris & Lycett, 1851, under Art. 70.3; Ampezzopleura tenuis Nutzel, 1998, fixed as type species of Ampezzopleura Bandel, 1991, under Art. 70.3; Proserpina nitida G. B. Sowerby II, 1839, designated type species of Despoena Newton, 1891; Buccinum glabratum Linnaeus, 1758, designated type species of Dipsaccus H. Adams & A. Adams, 1853; Murex ficus Linnaeus, 1758, designated type species of Ficula Swainson, 1835; Oncomelania hupensis Gredler, 1881, designated type species of Hemibia Heude, 1890; Murex metaxa Delle Chiaje, 1828, fixed as type species of Metaxia Monterosato, 1884 under Art. 70.3; Neridomus anglicus Cox & Arkell, 1950, fixed as type species of Neridomus J. Morris & Lycett, 1851, under Art. 70.3; Navicella clypeolum Recluz, 1843, designated type species of Orthopoma Gray, 1868; Trochus viadrinus M. Schmidt, 1905, fixed as type species of Parataphrus Chavan, 1954 under Art. 70.3; Helix pomatia Linnaeus, 1758, designated type species of Pentataenia A. Schmidt, 1855; Flammulina ponsonbyi Suter, 1897, fixed as type species of Phenacohelix Suter, 1892, under Art. 70.3; Cyrtolites corniculum Eichwald, 1860, fixed as type species of Pollicina Koken, 1895, under Art. 70.3; Purpurina elegantula d'Orbigny, 1850, designated as type species of Purpurina d'Orbigny, 1850, and lectotype of Turbo bellona d'Orbigny, 1850, designated as neotype of Purpurina elegantula; Pyramidella minuscula Monterosato, 1880, fixed as type species of Tiberia Jeffreys, 1884, under Art. 70.3; Cyclostoma delicatum Philippi, 1844, fixed as type species of Trachysma G. O. Sars, 1878, under Art. 70.3; Helix elegans Gmelin, 1791, fixed as type species of Trochoidea T. Brown, 1827, under Art. 70.3; Turritellopsis stimpsoni Dall, 1919, fixed as type species of Turritellopsis G. O. Sars, 1878, under Art. 70.3; Fusus averillii Gabb, 1864, fixed as type species of Volutoderma Gabb, 1876, under Art. 70.3; Voluta pepo Lightfoot, 1786, fixed as type species of Yetus Bowdich, 1822. Curnonidae d'Udekem d'Acoz, nom. nov., and Curnon d'Udekem d'Acoz, nom. nov., are established for Charcotiidae Odhner, 1926, and Charcotia Vayssiere, 1906, (between 27 March and 1 May), non Charcotia Chevreux, 1906 (January) [Amphipoda]; Yuopisthonematidae Nutzel, nom. nov., and Yuopisthonema Nutzel, nom. nov., are established for Opisthonematidae Yu, 1976, and Opisthonema Yu, 1974, non Gill, 1862 [Pisces]. The new family-group name Burnupiidae Albrecht is established in this work; and the names Scolodontina and Orthalicoidei are first used here to denote, respectively, a suborder containing the family Scolodontidae, and an infraorder containing the superfamily Orthalicoidea.

The nomenclature of Polyptychoceras, a Late Cretaceous heteromorph ammonite genus, and 18 related taxa is examined with a view to clarifying their nomenclatural availability, authors, dates of publication, and name-bearing types, on the basis of the International Code of Zoological Nomenclature (Fourth Edition). It is concluded that one family-group name, four genus-group names, and 12 species-group names are currently available. In contrast, the names Po. subundatum and Po. jimboi are unavailable. We also showed that Po. yubarense had been established by Shimizu (J Shanghai Sci Inst, Sect II, 1(11):159–226, 1935a).

Zoologica ScriptaVolume 36, Issue 6 p. 623-623 Free Access Corrigendum This article corrects the following: Notothrix halsei gen. n., sp. n., representative of a new family of freshwater cladocerans (Branchiopoda, Anomopoda) from SW Australia, with a discussion of ancestral traits and a preliminary molecular phylogeny of the order Kay Van Damme, Russell J. Shiel, H. J. Dumont, Volume 36Issue 5Zoologica Scripta pages: 465-487 First Published online: September 4, 2007 First published: 01 October 2007 https://doi.org/10.1111/j.1463-6409.2007.00304.xCitations: 2AboutSectionsPDF ToolsRequest permissionExport citationAdd to favoritesTrack citation ShareShare Give accessShare full text accessShare full-text accessPlease review our Terms and Conditions of Use and check box below to share full-text version of article.I have read and accept the Wiley Online Library Terms and Conditions of UseShareable LinkUse the link below to share a full-text version of this article with your friends and colleagues. Learn more.Copy URL Share a linkShare onFacebookTwitterLinked InRedditWechat Gondwanotrichidae nom. nov. pro Nototrichidae Van Damme, Shiel & Dumont, 2007 The authors (Van Damme et al.) of the paper ‘Notothrix halsei gen. n., sp. n., representative of a new family of freshwater cladocerans (Branchiopoda, Anomopoda) from SW Australia, with a discussion of ancestral traits and a preliminary molecular phylogeny of the order’ have alerted us to an error in their paper. The correction follows: In the previous issue of Zoologica Scripta [36(5): 465–487], we described a new monotypic family of Anomopoda (Crustacea: Branchiopoda) from SW Australia and named it as the Nototrichidae. However, the name of the type genus Notothrix Van Damme, Shiel & Dumont, 2007 turns out to be a junior homonym of the turbellarian genus Notothrix Hickman, 1955. Although the taxon described by Hickman (1955) was later synonymized with Umagilla Wahl, 1909 (Cannon 1987), this name remains available and therefore Notothrix Van Damme, Shiel & Dumont, 2007 is a junior homonym. As Notothrix Van Damme, Shiel & Dumont, 2007 is the type genus of the Nototrichidae Van Damme, Shiel & Dumont, 2007, the family name too becomes invalid, according to Article 39 of the International Code of Zoological Nomenclature (1999): ‘The name of a family group taxon is invalid if the name of its type genus is a junior homonym ...’. There are no synonyms for the type genus of the Nototrichidae. Therefore, we propose a new replacement name (nomen novum) for both the genus and the family. This is a strict nomenclatural issue that has further no effects on the content of the original paper. Family Gondwanotrichidae nom. nov. pro Nototrichidae Van Damme, Shiel & Dumont, 2007; diagnosis and description as in original publication. Type genus. Gondwanothrix nom. nov. pro Notothrix Van Damme, Shiel & Dumont, 2007 non Hickman, 1955. Type species. Notothrix halsei Van Damme, Shiel & Dumont, 2007. = Gondwanothrix halsei (Van Damme, Shiel & Dumont, 2007). Etymology. The name ‘Gondwanothrix’ consists of the prefix ‘Gondwano-’, referring to Gondwanaland, the supercontinent of the southern hemisphere that existed until its break-up in the Mesozoic, as we believe that this is one of the most primitive extant anomopods, dating back at least to that era (Van Damme et al. 2007). The word ‘thrix’ refers to the long setae on the posteroventral valve corner and the long spines on the postabdomen; the epitheton ‘halsei’ remains. Acknowledgements The authors wish to thank Steve Tracey and Dr Andrew Polaszek at the ICZN Secretariat for help and constructive advice. References Cannon, L. R. G. (1987). Two new rhabdocoel turbellarians, Umagilla pacifica sp. n. and U. karlingi sp. n. (Umagillidae), endosymbiotic with holothurians (Echinodermata) from the Great Barrier Reef and a discussion of sclerotic structures in the female system of the Umagillidae. Zoologica Scripta, 16 (4), 297– 303. Wiley Online LibraryWeb of Science®Google Scholar Hickman, V. V. (1955). Two new rhabdocoel turbellarians parasitic in Tasmania holothurians. Proceedings of the Royal Society of Tasmania, 89, 81– 97. Google Scholar International Commission on Zoological Nomenclature (1999). International Code of Zoological Nomenclature, 4th edn. Published by the International Trust for Zoological Nomenclature, c/o The Natural History Museum, London, UK. URL: http://www.iczn.org/iczn/index.jsp . Google Scholar Van Damme, K., Shiel, R. J. & Dumont, H. J. (2007). Notothrix halsei gen. n., sp. n., representative of a new family of freshwater cladocerans (Branchiopoda, Anomopoda) from SW Australia, with a discussion of ancestral traits and a preliminary molecular phylogeny of the order. Zoologica Scripta, 36 (5), 465– 487. Wiley Online LibraryWeb of Science®Google Scholar Citing Literature Volume36, Issue6November 2007Pages 623-623 ReferencesRelatedInformation

Corrigendum to “Assessing phylogenetic dependence of morphological traits using co-inertia prior to investigate character evolution in Loricariinae catfishes” Mol. Phylogenet. Evol. 46 (2008) 986–1002

The island of Madagascar is a renowned hotspot for adaptive radiations. Madagascar has been separated from mainland Africa since the end of the Jurassic, and from India since the Late Cretaceous. This long isolation, combined with the island’s large size and relatively few dispersal events has resulted in an avifauna characterized by a low species count and high endemism: for instance, 80% of the breeding Malagasy songbirds (Passeriformes) are endemic (Hawkins & Goodman 2003). A first series of papers (Cibois et al. 1999, 2001; Fjeldsa et al. 1999) on the phylogeny of the Malagasy taxa traditionally classified as Timaliidae, Sylviidae and Pycnonotidae (all families included in the large sylvioid clade) showed that several of these passerines form an original radiation endemic to the island. Because these results were based solely on a single kind of molecular marker (mitochondrial DNA sequences), the authors refrained at that time from giving a name to this clade. More recently, other studies using nuclear markers as well (Beresford et al. 2005; Johansson et al. 2008a, 2008b) confirm the existence of this Malagasy sylvioid radiation. The species that comprise this group exhibit a great variety of bill shapes, wing and tail proportions, and tarsus lengths. This diversity in morphology is linked to varieties of habitat and prey favoured by these insectivorous forest dwellers (Schulenberg 2003). Thus the endemic Malagasy sylvioid clade rivals other island radiations, including the vangas of Madagascar and the finches of the Galapagos, in ecological and morphological diversity. Several authors were inclined to consider this group at the family level, using the name ‘Bernieridae’. To our knowledge the first study using this name was the book “The natural history of Madagascar”, edited by S. M. Goodman and J. Benstead in 2003, where the name ‘Bernieridae’ appeared in two chapters (in Tingle et al. (2003: p. 522) and Hawkins & Goodman (2003: p. 1036), although Schulenberg (2003: p. 1131) referred to the Malagasy "warblers" in his chapter on the radiations of passerine birds on Madagascar). An alternative spelling for the family-group name, ‘Bernieriidae’, can be found in several personal pages on the internet, but we have not found an occurrence of this in any publication, as defined in the International Code of Zoological Nomenclature (4th edition, 1999). The name ‘Bernieridae’ was later used in several journal articles (Chouteau & Fenosoa 2008; Fuchs et al. 2008; Johansson et al. 2008a, 2008b), however, none of these have introduced the family-group name ‘Bernieridae’ according to the provisions of the International Code of Zoological Nomenclature, i.e. the nominal taxon was not explicitly indicated as intentionally new (Article 16.1) and the type genus was not cited (Article 16.2). In the present paper, we therefore propose to rectify this situation by correctly introducing the family-group name for the Malagasy sylvioid radiation.

For the first time, all family-group names in the superfamily Scarabaeoidea (Coleoptera) are evaluated using the International Code of Zoological Nomenclature to determine their availability and validity. A total of 383 family-group names were found to be available, and all are reviewed to scrutinize the correct spelling, author, date, nomenclatural availability and validity, and current classification status. Numerous corrections are given to various errors that are commonly perpetuated in the literature. A reversal of precedence is used to preserve the prevailing usage of the following family-group names: Eupariini Schmidt, 1910 (over Ataeniini Harold, 1868); Pachydemini Burmeister, 1855 (over Elaphocerini Blanchard, 1851); Heterosternina Bates, 1888 (over Macropnina Horn, 1866); and Anomalina Streubel, 1839 (over Euchlorina Hope, 1839).

The name, Galathea keijii, was given to a Miocene fossil species of squat lobster from Japan by Karasawa (1993: 39, pl. 6, figs. 1, 2, 3, 10). In the same year, the name, Galathea keijii, was established for a living species from the Andaman Sea and Arabian Sea (Tirmizi Javed 1993: 50, fig. 22). Therefore, both names are homonymous. The description of Karasawa was published in the Bulletin of the Mizunami Fossil Museum, no. 20, which, on the inside back cover states the date of publication as December 25, 1993. The description of Tirmizi Javed was published in "Indian Ocean galatheids (Crustacea: Anomura)" by the Marine Reference Collection and Resource Centre, University of Karachi. This work was published in 1993 but the day and month of publication were not cited in the publication. In the absence of evidence to the contrary, the date of publication for "Indian Ocean galatheids (Crustacea: Anomura)" is deemed to be the last day of 1993, by application of Article 21.3.2 of the International Code of Zoological Nomenclature (ICZN 1999: 20). As such, application of Article 52 on the principle of homonymy of the International Code of Zoological Nomenclature (ICZN, 1999: 56) shows that Galathea keijii Karasawa, 1993, takes precedence over Galathea keijii Tirmizi Javed, 1993. In accordance with Article 60 on the replacement of junior homonyms (ICZN 1999: 62, 63) a new replacement name, Galathea nasimae, is here proposed for Galathea keijii Tirmizi Javed, 1993. It is dedicated to the late Dr. Nasima M. Tirmizi (Marine Reference Collection and Resource Centre, University of Karachi).

Various problems posed by the 2012 Amendment to the International Code of Zoological Nomenclature are reviewed. They concern mainly the nomenclatural availability and the promulgation date of works published online and of the new nomina and nomenclatural acts they contain. A methodology is proposed for the analysis of these problems in many works published after 3 September 2012. A detailed survey of 120 herpetological online publications having nomenclatural implications is presented: 63 of them include unavailable nomina and nomenclatural acts, and 57 of them were made available through their printed versions at dates subsequent to that stated on their PDFs. Detailed proposals and recommendations to authors, editors and referees, publishers, libraries, and concerning the Code, Zoobank and the Commission, are offered to try to limit the negative impact of these problems in zootaxonomy.

Charles Johnson Maynard (1845–1929) was a self-educated naturalist, teacher, and dealer in natural history specimens and materials who conducted extensive field work throughout Florida, the Bahamas, and the Cayman Islands. He published prolifically on the fauna, flora, and anthropology of these areas. His publications included descriptions of 248 of the 587 validly proposed species-level taxa within Cerionidae, a family of terrestrial gastropods endemic to the islands of the tropical western Atlantic. After his death, his collection of Cerionidae was purchased jointly by the Museum of Comparative Zoology (MCZ) and the United States National Museum, with the presumed primary types remaining at the MCZ and the remainder of the collection divided between these two museums and a few other institutions. In this work, we provide 1) a revised collation of Maynard's publications dealing with Cerionidae, 2) a chronological listing of species-level taxa proposed in these works, 3) a determination of the number and status of name-bearing type specimens for each taxon within the context of the most recent edition of the International Code of Zoological Nomenclature, and 4) when necessary, the selection of a lectotype (for 185 taxa) or the designation of a neotype (for eight taxa) from among specimens in these museums to restrict the name-bearing type for each taxon to a single specimen, which is illustrated. Type material could not be located for three of the 248 species-level taxa proposed by Maynard. In these instances, neo-types are proposed to make these three taxa objective junior synonyms of other taxa. Strict application of the International Code of Zoological Nomenclature contradicts the authorship, publication date, and type status of name-bearing types for a significant number of Maynard's taxa that had been previously reported in the literature.

1.1.3 Nomenclatorial problems in usage of some family and genus names

Family-group names proposed for beetles belonging to the family Cerambycidae are catalogued and their availability is determined using the rules of the current International Code of Zoological Nomenclature. A synoptic classification of the family summarizes the validity of the names. Type genera of all family-group names are listed and the type species and stems of genera of available family-group names are included. A new family-group name, Elytracanthinini Bousquet (type genus: Elytracanthina Monn, 2005, a replacement name for Elytracantha Lane, 1955) is proposed for Elytracanthinae Lane, 1955. Ichthyosoma armatum Montrouzier, 1855 is designated as type species of Icthyosoma Boisduval, 1835. Reversal of precedence is used to preserve the validity of the following family-group names: Anaglyptides Lacordaire, 1868 (over Anaglyptisidae Gistel, 1848 [Buprestidae]); Dryobiini Arnett, 1962 (over Dryobiadae Gistel, 1856 [Ptinidae]); Hemilophitae Thomson, 1868 (over Amphionychitae Thomson, 1860) and Hétéropsides Lacordaire, 1869 (over Dichophyiaeidae Gistel, 1848). The following family-group names, although junior synonyms, are preserved as valid until an application is submitted to the International Commission on Zoological Nomenclature; in these cases a reversal of precedence could not be applied: Eurypodini Gahan, 1906 (over Zaracinae Pascoe, 1869); Macronides Lacordaire, 1868 (over Enchapteritae Thomson, 1861); Pyresthides Lacordaire, 1868 (over Pseudolepturitae Thomson, 1861 and Erythrinae Pascoe, 1866) and Stenoderinae Pascoe, 1867 (over Syllitae Thomson, 1864). A total of 238 valid cerambycid family-group names (413 available names) are recognized in the following 13 subfamilies: Vesperinae (1 valid family-group name), Oxypeltinae (1), Disteniinae (4), Anoplodermatinae (3), Philinae (1), Parandrinae (2), Prioninae (24), Spondylidinae (5), Necydalinae (1), Lepturinae (8), Lamiinae (80), Dorcasominae (1), and Cerambycinae (107).

Donegan (2012) proposed the family-group name "Ortalidaini n. tribe" for the chachalaca group in the avian family Cracidae (guans), and designated "Ortalis (or Ortalida)" [sic] Merrem, 1786, as the type genus. For reasons given below, the name "Ortalidaini" is showed to be misformed according to the International Code of Zoological Nomenclature (ICZN 1999), hereinafter the Code, and the correct stems of family-group names using Ortalis Merrem, 1786, as type genus are presented.

We are now over four decades into digitally managing the names of Earth's species. As the number of federating (i.e., software that brings together previously disparate projects under a common infrastructure, for example TaxonWorks) and aggregating (e.g., International Plant Name Index, Catalog of Life (CoL)) efforts increase, there remains an unmet need for both the migration forward of old data, and for the production of new, precise and comprehensive nomenclatural catalogs. Given this context, we provide an overview of how TaxonWorks seeks to contribute to this effort, and where it might evolve in the future. In TaxonWorks, when we talk about governed names and relationships, we mean it in the sense of existing international codes of nomenclature (e.g., the International Code of Zoological Nomenclature (ICZN)). More technically, nomenclature is defined as a set of objective assertions that describe the relationships between the names given to biological taxa and the rules that determine how those names are governed. It is critical to note that this is not the same thing as the relationship between a name and a biological entity, but rather nomenclature in TaxonWorks represents the details of the (governed) relationships between names. Rather than thinking of nomenclature as changing (a verb commonly used to express frustration with biological nomenclature), it is useful to think of nomenclature as a set of data points, which grows over time. For example, when synonymy happens, we do not erase the past, but rather record a new context for the name(s) in question. The biological concept changes, but the nomenclature (names) simply keeps adding up. Behind the scenes, nomenclature in TaxonWorks is represented by a set of nodes and edges, i.e., a mathematical graph, or network (e.g., Fig. 1). Most names (i.e., nodes in the network) are what TaxonWorks calls "protonyms," monomial epithets that are used to construct, for example, bionomial names (not to be confused with "protonym" sensu the ICZN). Protonyms are linked to other protonyms via relationships defined in NOMEN, an ontology that encodes governed rules of nomenclature. Within the system, all data, nodes and edges, can be cited, i.e., linked to a source and therefore anchored in time and tied to authorship, and annotated with a variety of annotation types (e.g., notes, confidence levels, tags). The actual building of the graphs is greatly simplified by multiple user-interfaces that allow scientists to review (e.g. Fig. 2), create, filter, and add to (again, not "change") the nomenclatural history. As in any complex knowledge-representation model, there are outlying scenarios, or edge cases that emerge, making certain human tasks more complex than others. TaxonWorks is no exception, it has limitations in terms of what and how some things can be represented. While many complex representations are hidden by simplified user-interfaces, some, for example, the handling of the ICZN's Family-group name, batch-loading of invalid relationships, and comparative syncing against external resources need more work to simplify the processes presently required to meet catalogers' needs. The depth at which TaxonWorks can capture nomenclature is only really valuable if it can be used by others. This is facilitated by the application programming interface (API) serving its data (https://api.taxonworks.org), serving text files, and by exports to standards like the emerging Catalog of Life Data Package. With reference to real-world problems, we illustrate different ways in which the API can be used, for example, as integrated into spreadsheets, through the use of command line scripts, and serve in the generation of public-facing websites. Behind all this effort are an increasing number of people recording help videos, developing documentation, and troubleshooting software and technical issues. Major contributions have come from developers at many skill levels, from high school to senior software engineers, illustrating that TaxonWorks leads in enabling both technical and domain-based contributions. The health and growth of this community is a key factor in TaxonWork's potential long-term impact in the effort to unify the names of Earth's species.

Is Myxomycetes (Amoebozoa) a Truly Ambiregnal Group? A Major Issue in Protist Nomenclature

In our recently published revised classification of the Icteridae (Remsen et al. 2016), we used the family group name Cassicinae Bonaparte, 1853 on the assumption that its type genus was Cassicus Illiger, 1811. We have since confirmed, after kind advice from Thomas Donegan (pers. comm. 2016), that Cassicus Illiger (1811: 214) is simply an unjustified emendation of Cacicus Lacépède, 1799, made clear by Illiger (l.c.) in a footnote to his description of Cassicus. Under Articles 32.5.3.2 and 35.4.2 of the International Code of Zoological Nomenclature (ICZN 1999), a family-group name formed from an unjustified emendation of the name of its type genus is to be corrected, unless the emendation has come into use as a substitute name or through prevailing use. This is not the case here, and accordingly we correct the spelling of Cassicinae to Cacicinae. Bonaparte (1853) retains authorship; and, although not expressly mentioned before, his originally ligatured suffix -eoe for the name was automatically corrected by us (Remsen et al. l.c.) under Articles 11.7.1.3 and 32.5.3 of the Code (ICZN l.c.).