Abstract
Pterosaurs are powered flight vertebrates that first emerged in the Late Triassic and persisted until the end of the Cretaceous. Recent studies on the ecological niches and habitats of pterosaurs, based on discoveries of Cretaceous bone beds in China, Brazil, Morocco, and other locations, suggest a decline in pterosaur diversity during the mid-Cretaceous. Various hypotheses have been proposed to interpret this phenomenon. This study focuses on newly discovered pterosaur tracks at the Upper Cretaceous (Truonian) Hwasun Seoyuri tracksite in Korea. The analysis is based on pterosaur tracks from in situ and ex situ specimens that originate from a variety of stratigraphic levels. The evidence found suggests that the Hwasun Seoyuri tracksite was occupied by small pterosaurs over an extended period, with indications of recurrent activity across at least six levels. Furthermore, the footprint sizes observed across these multiple layers follow a non-skewed normal distribution. Considering the known logarithmic growth characteristics of pterosaurs, this result suggests a predominantly immature population at this site. The prevalence of mid-Cretaceous non-marine pterosaur tracksites on the Korean Peninsula may reflect broader ecological shifts for pterosaurs during this period. This evidence is considered alongside the habitat shifts suggested by other non-marine pterosaur fossil sites in China and Brazil, as well as with indications of the decline in Late Cretaceous non-marine basins.
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