Abstract

Vertebrate tracks offer important paleobiological and paleoenvironmental insights as indicated in the increasing number of studies of modern and fossil tracks in recent decades (Laporte and Behrensmeyer, 1980; Loope, 1986; McKeever, 1991; Platt and Hasiotis, 2008; Milan et al., 2007; and references therein). Whereas fine-grained sediments typically preserve the greatest detail, a track record of many organisms is archived in sand-rich strata representing paleo-shorelines, river banks, and dunes (Frey and Pemberton, 1986; Lockley, 1986). Despite the dynamic nature and coarse texture of sandy media ( = substrates), many studies report abundant vertebrate and invertebrate tracks, from Paleozoic sandstones to Cenozoic depositional sequences (McKeever, 1991; Lea, 1996; Loope, 2006; Lucas et al., 2007). Aside from a few cursory investigations of vertebrate tracks in recent beach and dune environments, little research has focused on their distribution and preservation in unconsolidated sand-rich media (McKee, 1947; Loope, 1986; Lea, 1996). This is largely due to the difficulty of preserving the tracks and even greater challenge of detecting biogenic structures in coarse-grained sediments. New neoichnological research is underway that addresses this long-standing problem, making use of the combination of natural selective mineral sorting and the ability to image small-scale structures using geophysical techniques. In many coastal and eolian settings, such lithological anomalies as heavy-mineral concentrations (HMCs) represent density lag formation during episodes of increased wave or wind activity (Komar and Wang, 1984; Smith and Jackson, 1990; Buynevich et al., 2007; Mange and Wright, 2007). Such lithological anomalies not only act as environmental indicators, but also accentuate vertebrate and large invertebrate tracks due to their darker color (Fig. 1; …

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