FOOD AS A CONTROL OF A POPULATION OF WHITE-FOOTED MICE, PEROMYSCUS LEUCOPUS NOVEBORACENSIS (FISCHER)

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Mice were introduced to an island before and after food was supplied in excess of food consumed. The population of mice on the experimental island with food in excess is compared with populations of mice on the same island before the addition of food, on an adjacent island, and on the mainland. Before food was supplied in excess, death rate (death and dispersal) greatly exceeded birth rate and the population failed. After food was supplied in excess death rate of the second introduction decreased and population increased to greatly exceed the density of mice on the control island.The increased density of mice on the experimental island is the result of a decreased death rate, particularly in mice from birth to approximately 1 month of age. Increased birth rate may also be a factor. Food supply regulates the abundance of white-footed mice by affecting death rate and possibly birth rate. The most important effect of food supply is on the survival of young from birth to approximately 1 month of age.Evidence that intraspecific strife for space occurs in relatively dense population is presented. Intraspecific strife for space may regulate population of mice beyond the level of abundance determined by the supply of food.

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CitationsShowing 10 of 79 papers
  • Open Access Icon
  • Dissertation
  • Cite Count Icon 40
  • 10.31274/rtd-180813-7268
Habitat selection by small mammals of riparian communities: evaluating the effects of habitat alteration
  • Aug 3, 2015
  • Anthony Raymond Geier

Small mammals of riparian communities in Iowa were studied during the summer using liveand snap-trapping techniques. Six general habitat types were identified from the herbaceous vegetation on 28 study plots selected to represent a range of habitats from open fields to deciduous forest. Predominant habitat alterations were grazing, timber removal, and stream-channel realignment. Small-mammal species diversity was highest in channelized habitats and lowest in dry floodplains. An index of breadth of habitat usage was calculated for 9 species of mammals; white-footed mice (Peromyscus leucopus) exhibited the most generalized habitat usage. With the use of stepwise multiple regression, relationships were determined between small-mammal species abundances and 12 variables describing microhabitat features. In many instances, small-mammal numbers also were correlated significantly with each other. The potential effects of 6 habitat alterations on the 9 small-mammal species are predicted. J. WILDL. MANAGE. 44(1):16-24 Suitable habitat probably is the most important factor influencing the distribution and abundance of small mammals within their geographic ranges (Baker 1968:101, Vaughan 1972:250-256). Some small-mammal species have specific habitat requirements and consequently are limited in their distribution, whereas others occupy a wide variety of habitats (Kaufman and Fleharty 1974, Kirkland and Griffin 1974, Briese and Smith 1975, Miller and Getz 1976). Reports of the general habitats occupied by small mammals are common in the literature, but few studies have quantified the factors within a locality that influence a species abundance. Recently, small-mammal abundance and distribution have been related to several measures of habitat structure (M'Closkey 1975, M'Closkey and Fieldwick 1975, M'Closkey and Lajoie 1975, Dueser and Shugart 1978, Holbrook 1978). Habitat disturbances such as streamchannel realignment (Possardt and Dodge 1978), clear-cutting (Kirkland 1977, Martell and Radvanyi 1977), fire (Krefting and Ahlgren 1974, Fala 1975), and strip mining (Verts 1959, DeCapita and Bookhout 1975) can affect small-mammal populations and alter community composition. Regardless of the nature of disturbance, if vegetation is changed and habitat is altered, populations of some species may benefit while others are affected adversely. The objectives of our study were (a) to determine habitat preferences of some small mammals and the factors influencing their abundance; and (b) to quantify the effects of habitat alterations, particularly stream-channel realignment and grazing, on community composition and species abundance. Although this study was conducted in southwestern Iowa, the results are applicable to other riparian communities, especially those with similar small-mammal communities. We thank the people of Guthrie County for allowing us to sample small mammals on their property. A. Brackney and D. Schlapkohl assisted in the field, and 1 Journal Paper J-9463 of the Iowa Agriculture and Home Economics Experiment Station, Ames. Project 2085, funded by the U.S. Fish and Wildlife Service, Office of Biological Services, National Stream Alteration Team. 2 Present Address: 9025 W. Herbert, Milwaukee, WI 53225. 16 J. Wildl. Manage. 44(1):1980 This content downloaded from 157.55.39.243 on Thu, 06 Oct 2016 04:44:14 UTC All use subject to http://about.jstor.org/terms HABITAT SELECTION BY SMALL MAMMALS * Geier and Best 17 D. F. Cox and M. Hand provided help with statistical analyses. Vegetation data were collected by J. P. Vogler. R. B. Dahlgren, W. L. Franklin, and R. Q. Landers reviewed an earlier draft of the

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  • Research Article
  • Cite Count Icon 27
  • 10.1111/j.1558-5646.1965.tb03324.x
ISLAND POPULATIONS AND GENE FLOW IN THE DEER MOUSE,PEROMYSCUS LEUCOPUS
  • Dec 1, 1965
  • Evolution
  • Walter Sheppe

EvolutionVolume 19, Issue 4 p. 480-495 ArticleFree Access ISLAND POPULATIONS AND GENE FLOW IN THE DEER MOUSE, PEROMYSCUS LEUCOPUS Walter Sheppe, Walter Sheppe Department of Biology, State University of New York, BuffaloSearch for more papers by this author Walter Sheppe, Walter Sheppe Department of Biology, State University of New York, BuffaloSearch for more papers by this author First published: December 1965 https://doi.org/10.1111/j.1558-5646.1965.tb03324.xCitations: 6AboutPDF ToolsRequest permissionExport citationAdd to favoritesTrack citation ShareShare Give accessShare full text accessShare full-text accessPlease review our Terms and Conditions of Use and check box below to share full-text version of article.I have read and accept the Wiley Online Library Terms and Conditions of UseShareable LinkUse the link below to share a full-text version of this article with your friends and colleagues. Learn more.Copy URL Share a linkShare onFacebookTwitterLinked InRedditWechat Citing Literature Volume19, Issue4December 1965Pages 480-495 ReferencesRelatedInformation

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  • Cite Count Icon 40
  • 10.1139/z77-220
Energy requirements during reproduction of Peromyscus maniculatus
  • Oct 1, 1977
  • Canadian Journal of Zoology
  • Lucius L Stebbins

Weight-specific food consumption, gross daily energy consumption, and weights of breeding groups, lactating females, and young litters were monitored between time of mating and 2 weeks after weaning. No observable changes in weight-specific or gross daily energy consumption was evident in the 3 weeks before the week in which birth occurred, although weights of females had by then increased an average of 36%. Gross daily energy consumption increased by 96%, 136%, and 194%, respectively, during the 1st, 2nd, and 3rd weeks after birth. Weights of females dropped significantly in the week before weaning.

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  • Research Article
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  • 10.1002/ecs2.2972
Tick control bait box use by Peromyscus spp. influenced by habitat placement but raises questions on disease ecology
  • Dec 1, 2019
  • Ecosphere
  • Erika T Machtinger + 1 more

Abstract White‐footed mice, Peromyscus leucopus (Rafinesque), and deer mice, Peromyscus maniculatus Gloger, are considered important reservoir hosts for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Host‐targeted and nonlethal Select TCS bait boxes have been shown to be effective at killing ticks by delivering small doses of fipronil to wild Peromyscus spp., attracted to the provided bait. This results in reductions in the tick Ixodes scapularis, the vector of Borrelia burgdorferi, in the environment. However, habitat influences on bait box use, interactions with bait boxes, and overall use of bait by small mammals have not been evaluated. These factors may influence the use of this tick control measure. In the current study, bait box use by small mammals was evaluated at five field sites in Maryland, USA, during a two‐year study period. Bait consumption in relation to specific described habitats evaluated at each site was recorded, and bait nutrition was compared to nutrition from common northeastern acorn species. Rodent behavior around bait boxes was analyzed using camera traps as well. Bait boxes were used more frequently in herbaceous and shrub habitats than those placed in areas with bare ground or canopy cover alone. Significant amounts of bait were consumed throughout the study, but in some habitat types bait boxes were not used. The bait was found to be similar to native acorns in energy content, suggesting that it may serve as a supplement for natural foods which may have population dynamic implications. Behavioral analysis suggested that Peromyscus spp. foraged around bait boxes regardless of bait presence, but they did not forage in unbaited boxes as frequently. This may suggest the use of odors alone could be enough for attracting rodents to bait boxes potentially reducing the negative effects of baiting while maintaining protective tick control measures.

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  • Cite Count Icon 68
  • 10.1890/07-1935.1
PARASITES PREVENT SUMMER BREEDING IN WHITE-FOOTED MICE,PEROMYSCUS LEUCOPUS
  • Aug 1, 2008
  • Ecology
  • Kurt J Vandegrift + 2 more

Food and parasites can independently play a role in destabilizing population fluctuations of animals, and yet, more than 50 years ago, David Lack proposed that these two factors should act in concert. We examined the role of these factors on the vital rates of free-living white-footed mice (Peromyscus leucopus) over the summer and autumn months. We used a replicated factorial experiment in which deer exclosures doubled acorn availability and anthelmintic application reduced gastrointestinal helminths. Specifically, we wanted to know if either factor or an interaction between the two accounted for the midsummer breeding hiatus observed in this species. We found no influence of habitat quality on mouse breeding, vital rates, or demography; however, anthelmintic treatment resulted in mice continuing to reproduce during the hiatus at the same rate as previously, and they also exhibited increased body condition, growth rate, and survival. These results provide evidence that gastrointestinal helminths reduce P. leucopus reproductive output in central Pennsylvania, and these effects on reproduction could play a role in the unstable dynamics of small mammals.

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  • 10.1016/j.cropro.2003.08.005
Influence of a granivorous diversionary food on population dynamics of montane voles ( Microtus montanus), deer mice ( Peromyscus maniculatus), and western harvest mice ( Reithrodontomys megalotis)
  • Oct 21, 2003
  • Crop Protection
  • Thomas P Sullivan + 1 more

Influence of a granivorous diversionary food on population dynamics of montane voles ( Microtus montanus), deer mice ( Peromyscus maniculatus), and western harvest mice ( Reithrodontomys megalotis)

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  • Cite Count Icon 28
  • 10.2307/1932672
Effects of Chlorpromazine on Aggression in Laboratory Populations of Wild House Mice
  • May 1, 1967
  • Ecology
  • Stephen Vessey

Three confined, freely growing populations of wild house mice (Mus musculus) were studies to determine possible functions of aggressive behavior in regulating population size. Effects of a tranquilizer, chlorpromazine, on aggression and population parameters were measured. In singly caged pairs of mice chlorpromazine, mixed with the food at a rate of 0.75 mg/g food, significantly increased the number of days between litters. Also, significantly fewer young were born and weaned when compared with pairs given a control diet. Thus chlorpromazine produced no direct enhancement of reproduction. At the same dose chlorpromazine reduced fighting among C57 male mice. The growth curves of the populations prior to treatment were generally sigmoid, upper limits varying in each population. Birth and survival rates of infants declined with increasing population size. Mortality rates of adults did not vary significantly with population size. Due to a higher mortality of post—weaned males than females, sex ratios generally favored females. Aggressive rates (interactions per min) increased as the population increased, then leveled off after the after the population reaches a peak. When adjusted for population size, the rates followed the same pattern. Adult males established territories in all three cages, up to seven such males being present in a cage at one time. These males significantly heavier than non—territorial males of similar age and initiated up to 8% of the fights. Adrenal glands of territorial males were significantly lighter and thymuses significantly heavier than those of non—territorial males. Females did not defend territories but accounted for up to 50% of the fighting. Chlorpromazine (0.75 mg) was given to one population for 111 days and to a second population for 60 days after each had leveled off. Renewed population growth occurred during drug administration and aggressive rates were markedly reduced. The only effect of the drug on measured population parameters was increased infant survival. When the drug was removed from one population the numbers of mice declined and the aggressive rate increased. Reduced aggression must have produced its effect on population size through some factor(s) influencing infant survival.

  • Research Article
  • Cite Count Icon 90
  • 10.2307/1934745
Field Populations of Deermice with Supplemental Food
  • Jan 1, 1971
  • Ecology
  • R A Fordham

A description is given of a field experiment from winter through fall in which excess artificial food was provided in some areas with Peromyscus maniculatus, but not in others. Population size and production of young increases, and adult but not juvenile survival improves slightly with additional food; hence the hypothesis that no differences in breeding and survival would exist between experimental and control areas is rejected in part. The numbers of adult males in experimental and control situations are similar, suggesting that the male portion of the breeding population may, as hypothesized by Sadleir (1965) and Healey (1967), be held relatively constant during breeding by behavioral mechanisms. Attention is drawn to the different numerical responses of males and females, and the possibility of different regulating mechanisms for the sexes.

  • Research Article
  • Cite Count Icon 13
  • 10.2307/2425464
Food as a Limiting Factor and Selective Agent for Genic Heterozygosity in the Cotton Mouse Peromyscus gossypinus
  • Jul 1, 1984
  • American Midland Naturalist
  • Michael W Smith + 2 more

Supplemental food was added to six of 12 1-ha grids for 14 months. At the end of this time, there were approximately 3.5 times as many cotton mice on the areas receiving food than on the control areas. The direct and/or indirect effects of food was limiting to population number on the control areas, and its lower availability apparently selected for increased heterozygosity at the esterase-1 (EST-1) locus. EST-1 may be an indicator of overall heterozygosity and animals which are more heterozygous may be better competitors for the limited food resource. The lack of significant effects on the other two variable loci and the slight possibility of a null allele at EST-1 lowers our confidence in this hypothesis. However, some sort of selection was operating in these populations. 43 references, 1 figure, 2 tables.

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  • Research Article
  • Cite Count Icon 23
  • 10.1242/jeb.210963
Developmental delay in shivering limits thermogenic capacity in juvenile high-altitude deer mice (Peromyscus maniculatus)
  • Jan 1, 2019
  • Journal of Experimental Biology
  • Cayleih E Robertson + 1 more

Many endotherms native to cold and hypoxic high-altitude (HA) environments have evolved a highly vascularized and aerobic skeletal muscle. This specialized muscle phenotype contributes via shivering to an enhanced capacity for aerobic thermogenesis (cold-induced V̇ O2,max). However, it is unclear how selection at HA for shivering thermogenesis acts early in the development of small altricial mammals, which are born with immature skeletal muscles and without the capacity for homeothermic endothermy. We have previously shown that postnatal maturation of brown adipose tissue and non-shivering thermogenesis is delayed in HA native deer mouse pups (Peromyscus maniculatus). To assess whether HA adaptation has also altered the developmental program of skeletal muscle and shivering thermogenesis, we used laboratory-reared descendants of deer mice native to low altitude (LA, 430 m a.s.l.) and HA (4350 m a.s.l.) and a LA congeneric outgroup (P. leucopus). We found that LA juveniles were able to shiver robustly at 2 weeks after birth. However, HA juveniles were unlikely able to shiver at this point, resulting in a 30% lower capacity for thermoregulation compared with lowlanders. It was only at 27 days after birth that HA juveniles had established the aerobic muscle phenotype characteristic of HA adults and a superior cold-induced V̇ O2,max compared with LA mice of the same age. The capacity for shivering may be delayed in HA mice to allow energy to be allocated to other important processes such as growth.

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  • Oecologia
  • J�Rgen Jacobs

1. In the Klostersee in southern Federal Republic of Germany, Daphnia hyalina (H) and D. cucullata (C) have coexisted for at least 50 years. in 1971/72 they exhibited seasonal density fluctuations which are in accord with predictions from a model based on alternative adaptations of two competitors to natality (food utilization) and mortality (escape from enemies) in an environment that fluctuates with respect to food and intensity of predation. 2. In general, both species fluctuate in a similar fashion. Birth (b), death (m) and growth rates (r) are correlated between both species. Furthermore, in either species birth and death rates are positively correlated. But the birth and death rates of H are about 1.7 times as large as those of C, resulting in stronger density fluctuations of H. This fact, in combination with the seasonal course of the b/m-ratio produces a marked seasonal segregation of the abundances of both species, with H dominating in early summer and C dominating in autumn and winter. 3. In both species, the fluctuations of the growth rate are mainly determined by fluctuations of the death and not the birth rate. 4. Birth and death rates are much higher in the warm season than in the cold season, but births dominate in the cold, deaths in the warm season. Evidence is presented that mortality is mainly due to predation by fish. 5. The differences of natality and mortality which cause the seasonal segregation between the two species, exist independent of seasonal temperature fluctuations. But, there is an additional temperature effect accentuating the differences. 6. A partial correlation analysis shows that by measuring the density of one species and lake temperature, the density difference between both species and, hence, the density of the second species can be predicted with good precision. 7. The importance of the observed temporal segregation and other factors for the stabilized coexistence of both species is discussed.

  • Research Article
  • Cite Count Icon 10
  • 10.1086/220486
The Theory of Population Growth Cycles
  • Sep 1, 1949
  • American Journal of Sociology
  • Donald Olen Cowgill

Four theoretical types of population growth cycles are presented. Growth cycles from lower death rates in combination with stationary birth rates seem to have characterized preindustrial societies. A growth cycle resulting from a lag of the birth rate behind the death rate while both are declining appears to have accompanied industrialization. Future growth cycles in industrialized societies will probably depend upon increased birth rates in combination with "sticky" death rates. The fourth theoretical cycle in which death rates lag behind birth rates while both increase has neither precedent nor prospect.

  • Research Article
  • 10.24147/2312-1300.2022.9(4).127-142
Этнодемографическая характеристика населения Чукотки
  • Jan 1, 2022
  • Herald of Omsk University. Series: Historical studies
  • O P Kolomiets

The article presents an ethno-demographic characteristic of the population of Chukotka from the 18th century to the present. Until now. The source base of the study is statistical materials, published documents concerning the number and characteristics of settlement, the way of life of the indigenous people of Chukotka, scientific literature on the problems of ethnic demography. The paper presents a comparative analysis of the results of surveys and censuses of the population of Chukotka for 1897, 1926/27, 1939, 1959, 1970, 1979, 1989, 2002, 2010 (and partly 2020) The main components of the change in the population and the national composition of the region (natural, general, migration population growth) in the Soviet and post-Soviet periods are characterized. According to the data of the 2002 and 2010 censuses, the number of Chukchi, Koryaks, and Yukagirs increased during the intercensal period; the number of Eskimos, Evens, Chuvans and Kereks decreased. The ambiguous demographic picture among indigenous peoples is connected, on the one hand, with the real processes of depopulation in some of them, and, on the other hand, with the qualitative side of the census. The main reasons for the decline in the number of indigenous people of Chukotka: problems with natural growth against the background of high birth and death rates. The decline in the birth rate is associated with a change in traditional attitudes towards having many children, a decrease in marriage rates and an increase in the proportion of single-parent families. Although the Chukchi, Eskimos, Evens and Yukagirs are classified by demographers as a group with an increased birth rate. The demographic situation is also influenced by the increase in the number of mixed marriages. It should be noted that despite the decline in the birth rate among the indigenous inhabitants of Chukotka, its rates are still higher than among the visiting population of the region. The increase in mortality rates in Chukotka is due to a change in the age structure of the inhabitants, since in the 1990s a significant part of the able-bodied population left the district, the growth in the share of indigenous people in the structure of total mortality increased. The life expectancy of indigenous peoples is significantly lower than the national and regional indicators. The current ethno-demographic situation in Chukotka will become clear after the publication of the results of the 2020 All-Russian Population Census. It can be stated with confidence that there is a significant decrease in the number of permanent residents of the region, and the number of indigenous people in the total population of the ChAO continues to increase.

  • Research Article
  • Cite Count Icon 827
  • 10.1214/aoms/1177730285
On the Generalized "Birth-and-Death" Process
  • Mar 1, 1948
  • The Annals of Mathematical Statistics
  • David G Kendall

The importance of stochastic processes in relation to problems of population growth was pointed out by W. Feller [1] in 1939. He considered among other examples the "birth-and-death" process in which the expected birth and death rates (per head of population per unit of time) were constants, $\lambda_o$ and $\mu_o$, say. In this paper, I shall give the complete solution of the equations governing the generalised birth-and-death process in which the birth and death rates $\lambda(t)$ and $\mu(t)$ may be any specified functions of the time. The mathematical method employed starts from M. S. Bartlett's idea of replacing the differential-difference equations for the distribution of the population size by a partial differential equation for its generating function. For an account of this technique,$^1$ reference may be made to Bartlett's North Carolina lectures [2]. The formulae obtained lead to an expression for the probability of the ultimate extinction of the population, and to the necessary and sufficient condition for a birth-and-death process to be of "transient" type. For transient processes the distribution of the cumulative population is also considered, but here in general it is not found possible to do more than evaluate its mean and variance as functions of $t$, although a complete solution (including the determination of the asymptotic form of the distribution as $t$ tends to infinity) is obtained for the simple process in which the birth and death rates are independent of the time. It is shown that a birth-and-death process can be constructed to give an expected population size $\bar n_t$ which is any desired function of the time $t$, and among the many possible solutions the unique one is determined which makes the fluctuation, Var$(n_t)$, a minimum for all. The general theory is illustrated with reference of two examples. The first of these is the $(\lambda_0, \mu_1t)$ process introduced by N. Arley [3] in his study of the cascade showers associated with cosmic radiation; here the birth rate is constant and the death rate is a constant multiple of the "age, $t$, of the process. The $\bar n_t$-curve is then Gaussian in form, and the process is always of transient type. The second example is provided by the family of "periodic" processes, in which the birth and death rates are periodic functions of the time $t$. These appear well adapted to describe the response of population growth (or epidemic spread) to the influence of the seasons.

  • Research Article
  • Cite Count Icon 78
  • 10.4319/lo.1992.37.1.0001
Population dynamics and body‐size selection in Daphnia
  • Jan 1, 1992
  • Limnology and Oceanography
  • Alan J Tessier + 2 more

We examined the population dynamics and seasonal change in body size of a population of Daphnia galeata mendotae experiencing strong vertebrate predation. The intensity of food limitation was quantified seasonally by in situ food addition. We also collected live animals early and late in the summer for laboratory studies designed to estimate genetic variation and seasonal change in life‐history traits for the population.The population was characterized by nearly equal short‐term birth and death rates that were strongly correlated with population density, suggesting density dependence. The population generally exhibited very high birth and death rates and no significant response to experimental food addition. The one exception was June 1988, when the population achieved high densities and exhibited strong food limitation. Throughout both summers, minimal and average body sizes of adults decreased.Laboratory studies document substantial genetic variation for body size and other life‐history traits among animals collected early in summer. By late summer, however, there was little genetic variation for these same traits. Furthermore, clones collected early in summer were genetically larger in body size at birth and maturity than clones collected in late summer. We interpret these results as indicating natural selection for body size.

  • Research Article
  • Cite Count Icon 4
  • 10.1103/physreve.109.024224
Catastrophic and noncatastrophic population crashes in a bitrophic system with dynamic additional food provision to cooperative predators.
  • Feb 28, 2024
  • Physical Review E
  • Saswati Biswas + 2 more

In this article we contemplate the dynamics of an additional food-provided prey-predator system. We assume that the behavior of cooperative predators induces fear in prey, which radically affects the prey's birth and death rates. We observe that the structural instability imposed by strong cooperative hunting among predators goes away with higher intensities of fear levels affecting the prey's reproductive output and mortality. High levels of prey refuge are not conducive to the survival of predators. In such a situation, adequate supply of high-quality additional food is favorable regarding the persistence and stability of the system. Interestingly, the system potentially exhibits two stable configurations under identical ecological conditions by allowing different bifurcation scenarios, including saddle-node and backward bifurcations, and associated hysteresis effects with prey refuge along with additional food quantity and quality. In the stochastic environment, the system experiences critical transitions through bifurcation-induced tipping events with time-varying additional food for predators. Enhanced disturbance events promote noise-induced switching and tipping events. Finally, our investigation explores whether impending population crashes resulting from the variability of additional food quantity and quality can reliably be predicted using early warning signals in the context of redshifted noise. Overall, our results may provide insights for finding control strategies in the context of community ecology.

  • Research Article
  • Cite Count Icon 572
  • 10.1086/282748
The Concept of r- and K-Selection: Evidence from Wild Flowers and Some Theoretical Considerations
  • Jan 1, 1972
  • The American Naturalist
  • Madhav Gadgil + 1 more

The relationship of the demographic parameters of a population to its ecological niche constitutes one of the central problems of population biology. A most interesting theoretical notion pertinent to this problem is r- and K-selection (MacArthur 1962; Cody 1966; MacArthur and Wilson 1967; Hairston, Tinkle, and Wilbur 1970; Roughgarden 1971). The central idea of r- and K-selection is that populations living in environments imposing high density-independent (D.I.) mortality (r-strategists) will be selectively favored to allocate a greater proportion of resources to reproductive activities at the cost of their capabilities to propagate under crowded conditions, and conversely, populations living in environments imposing high density-dependent (D.D.) regulation (K-strategists) will be selectively favored to allocate a greater proportion of resources to nonreproductive activities, at the cost of their capabilities to propagate under conditions of high D.I. mortality. From the argument just stated, it may be deduced that the birth rate of an r-strategist will be greater than that of a related K-strategist. However, increased birth rate under conditions of high D.I. mortality is not sufficient evidence for an r-strategy, because, as demonstrated later in this paper, any increase in D.I. mortality must by itself produce a new equilibrium of birth and death rates at higher values of both. The crucial evidence needed for r- and K-selection is whether an organism is allocating a greater proportion of its resources to reproductive activities (r-strategists) than another related one (K-strategist) under any and all D.D. and D.I. mortality conditions.

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