Abstract
of Tehuantepec and eastern Panama was and always had been devoid of primary (obligatory) freshwater fishes prior to the very late Tertiary. Instead, secondary freshwater fishes evolved in this area during the Neogene and perhaps for a longer period, the Poeciliidae having probably had a longer history within the area than the Cichlidae. In the late Tertiary a very few North American immigrants entered the area from the north, and towards the end of the Pliocene the closing of the very ancient Panama sea gap permitted an influx of South American primary types. Most of these have not yet gotten past Costa Rica, but a few aggressive characids have reached Guatemala or southern Mexico and one has
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