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Cuttlefish Turn Slowly but Tightly with Directional Flexibility Using Short Vortex Ring Jets.

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Abstract Cephalopods, such as squids and cuttlefishes, play an integral role in food web dynamics as both prey and predator, whereby proficient turning is required for successful predator avoidance and prey capture. However, surprisingly little is known about the turning capabilities of most cephalopods. In this study, body movements and 3D flow fields around adult dwarf cuttlefish, Sepia bandensis, were recorded during a range of maneuvers to quantify turning performance. While significant variation in turning capabilities was observed, S. bandensis, on average, turned tightly (mean length-specific turning radius = 0.14±0.013 [SEM]; minimum length-specific turning radius = 0.013±0.002) but relatively slowly (average angular velocity = 45.85°±2.70° s-1; maximum angular velocity = 110.34°±7.09° s-1). Jet properties were not reliable predictors of turn performance, as S. bandensis relied primarily on short vortex ring jets (length-to-diameter ratio of jet vorticity = 2.47±0.18) of moderate velocity (average jet velocity ∼ 14 cm s-1; maximum jet velocity ∼ 22 cm s-1) irrespective of turn type. The orientation of the turn (arms-first vs. tail-first) did not have a significant effect on kinematic or hydrodynamic properties, and most turns were performed arms-first (72.6% of turns). Compared to other cephalopods, cuttlefish turned using shorter jets with similar velocity profiles. These results are consistent with the residence of S. bandensis in complex benthic habitats, where tight, controlled turns facilitated by short jet pulses and high turning proficiency in either orientation are needed.

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  • Research Article
  • Cite Count Icon 50
  • 10.2118/1288-g
Design and Operation of Jet-Bit Programs For Maximum Hydraulic Horsepower, Impact Force or Jet Velocity
  • Dec 1, 1960
  • Transactions of the AIME
  • H.A Kendall + 1 more

Published in Petroleum Transactions, AIME, Volume 219, 1960, pages 238–250. Abstract Several investigations in recent years have shown that drilling rates are increased significantly with increased hydraulic horsepower. But, there has been no overall method of designing jet-bit programs that efficiently uses the surface power. A study of present practices indicates that frequently as little as 50 per cent of the possible effects at the bit are used. Some observers have indicated that the best utilization of hydraulic horsepower (maximum effect on drilling rate) occurs when the bit hydraulic horsepower is maximum; others have stated that jet impact force is more important, and others have believed that maximum jet velocity is required. Limited efforts to date have shown some optimum conditions for bit hydraulic horsepower and impact, but these conditions cannot exist during drilling of a large part of the hole and do not provide a basis for designing a complete jet-bit program. This paper shows the maximum obtainable bit horsepower, impact force and jet velocity at all depths, taking into account the limitations of the pump, piping, hole and minimum circulating rate for adequate cuttings removal. Ranges of operation are developed; and flow rates, surface pressure and bit pressures are specified for each range to provide a maximum of any one of the desired effects. It also is shown that, by proper selection of nozzle sizes and by following the rules presented, the maximum obtainable quantities can be effectively utilized from surface to total depth. Finally, a simple graphical method of selecting nozzle sizes and flow rates is presented which can be used with familiar bit-company hydraulic tables and calculators to design jet-bit programs for maximum bit hydraulic horsepower, impact or jet velocity, as desired. These programs make most effective use of the pumps. Heretofore, there was no method available for designing field tests which adequately separated the effects of bit horsepower, impact and jet velocity. The programs and procedures developed in the paper are dissimilar and, when used in future field testing, should demonstrate which program is the most important in obtaining the fastest drilling rate.

  • Research Article
  • 10.1097/00005768-200305001-01792
RELATIONSHIPS AMONG UPPER BODY STRENGTH PARAMETERS AND ANGULAR VELOCITIES IN COMPETITIVE FIGURE SKATERS
  • May 1, 2003
  • Medicine & Science in Sports & Exercise
  • S M Macdonnell + 2 more

The ability to complete triple and quadruple jumps in figure skating is increasingly determining competitive outcomes. A closed arm position during rotating jumps increases the angular velocity and spin rate. Therefore, upper body strength may contribute to a successful jump. PURPOSE To examine the correlation between upper body strength, power and body size with off-ice vertical jump height (JH) and off-ice angular velocities. METHODS Seventeen competitive female figure skaters were tested for shoulder adduction/abduction and flexion/extension strength and power at speeds of 300, 400, and 450 degrees/second. Peak torque, torque as a percentage of body weight, and average power were measured for all of the isokinetic tests. Off-ice angular velocities were calculated using high-speed, three-dimensional cameras. Trials were filmed at 180 frames per second for 3 seconds. Subjects took a 2-step approach, jumped, and spun attempting to complete a maximal number of rotations. Measurements included: maximal angular velocity (MAV), (JH), time in the air (TIA), revolutions in the air (RIA), and average angular velocity (AAV). Correlations were examined by a Pearson Product correlation matrix. RESULTS No significant correlations were identified between the strength or power measurements and any of the angular velocities. However, off-ice MAV and AAV are only 78 and 80% of on-ice angular velocities. Both JH and TIA were correlated with RIA (P ≤.01). Age and BMI were also found to be inversely correlated with RIA respectively (P ≤.01); (P ≤ .05). CONCLUSION Although the results do not indicate a relationship between upper body strength and power and MAV or AAV, it is thought that the off—ice spin rates were too slow to adequately evaluate this relationship. Results do indicate a need to maintain a compact body shape, which may partially explain the success of younger, smaller female skaters. Results also indicate that body shape (> BMI) will change with age-related developmental changes, which will lead to increased torso inertia.

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  • Cite Count Icon 38
  • 10.1016/s0065-2504(08)60030-6
Generalist Predators, Interaction Strength and Food-web Stability
  • Jan 1, 1999
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Generalist Predators, Interaction Strength and Food-web Stability

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  • 10.4028/www.scientific.net/amm.187.63
Visualization of Supersonic Non-Newtonian Liquid Jets
  • Jun 1, 2012
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  • Anirut Matthujak + 3 more

This paper describes the characteristics of supersonic non-Newtonian liquid jets injected in ambient air. The main focus is to visualize three types of time-independent non-Newtonian liquid jet and to describe their behaviors. Moreover, comparisons between their dynamic behaviors with Newtonian liquid jet are reported. The supersonic liquid jets are generated by impact driven method in a horizontal single-stage power gun. Jets have been visualized by the high speed digital video camera and shadowgraph method. Effects of different liquid types on the jet penetration distance, average jet velocity and other characteristics have been examined. From shadowgraph images, the unique dynamic behaviors of each non-Newtonian liquid jets are observed and found obviously different from that of the Newtonian liquid jet. The maximum average jet velocity of 1,802.18 m/s (Mach no. 5.30) has been obtained. The jet penetration distance and average velocity are significantly varied when the liquid types are different.

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Jet flow in steadily swimming adult squid
  • Mar 15, 2005
  • Journal of Experimental Biology
  • Erik J Anderson + 1 more

Although various hydrodynamic models have been used in past analyses of squid jet propulsion, no previous investigations have definitively determined the fluid structure of the jets of steadily swimming squid. In addition, few accurate measurements of jet velocity and other jet parameters in squid have been reported. We used digital particle imaging velocimetry (DPIV) to visualize the jet flow of adult long-finned squid Loligo pealei (mantle length, L(m)=27.1+/-3.0 cm, mean +/-S.D.) swimming in a flume over a wide range of speeds (10.1-59.3 cm s(-1), i.e. 0.33-2.06 L(m) s(-1)). Qualitatively, squid jets were periodic, steady, and prolonged emissions of fluid that exhibited an elongated core of high speed flow. The development of a leading vortex ring common to jets emitted from pipes into still water often appeared to be diminished and delayed. We were able to mimic this effect in jets produced by a piston and pipe arrangement aligned with a uniform background flow. As in continuous jets, squid jets showed evidence of the growth of instability waves in the jet shear layer followed by the breakup of the jet into packets of vorticity of varying degrees of coherence. These ranged from apparent chains of short-lived vortex rings to turbulent plumes. There was some evidence of the complete roll-up of a handful of shorter jets into single vortex rings, but steady propulsion by individual vortex ring puffs was never observed. Quantitatively, the length of the jet structure in the visualized field of view, L(j), was observed to be 7.2-25.6 cm, and jet plug lengths, L, were estimated to be 4.4-49.4 cm using average jet velocity and jet period. These lengths and an average jet orifice diameter, D, of 0.8 cm were used to calculate the ratios L(j)/D and L/D, which ranged from 9.0 to 32.0 and 5.5 to 61.8, respectively. Jets emitted from pipes in the presence of a background flow suggested that the ratio between the background flow velocity and the jet velocity was more important than L/D to predict jet structure. Average jet velocities in steadily swimming squid ranged from 19.9 to 85.8 cm s(-1) (0.90-2.98 L(m) s(-1)) and were always greater in magnitude than swimming speed. Maximum instantaneous fluid speeds within squid jets ranged from 25.6 to 136.4 cm s(-1). Average jet thrust determined both from jet velocity and from three-dimensional approximations of momentum change in successive jet visualizations showed some differences and ranged from 0.009 to 0.045 N over the range of swimming speeds observed. The fraction by which the average jet velocity exceeded the swimming speed, or 'slip', decreased with increasing swimming speed, which reveals higher jet propulsive efficiency at higher swimming speeds. Jet angle, subtended from the horizontal, decreased from approximately 29 degrees to 7 degrees with increasing swimming speed. Jet frequency ranged from 0.6 to 1.3 Hz in the majority of swimming sequences, and the data suggest higher frequencies at the lowest and highest speeds. Jet velocity, angle, period and frequency exhibited increased variability at speeds between 0.6 and 1.4 L(m) s(-1). This suggests that at medium speeds squid enjoy an increased flexibility in the locomotive strategies they use to control their dynamic balance.

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  • Cite Count Icon 51
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Doppler echocardiographic findings in adults with severe symptomatic valvular aortic stenosis
  • Dec 1, 1991
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Resource tracking in marine parasites: going with the flow?
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Understanding how diversity interacts with energy supply is of broad ecological interest. Most studies to date have investigated patterns within trophic levels, reflecting a lack of food webs which include information on energy flow. We added parasites to a published marine energy‐flow food web, to explore whether parasite diversity is correlated with energy flow to host taxa. Parasite diversity was high with 36 parasite taxa affecting 40 of the 51 animal taxa. Adding parasites increased the number of trophic links per species, trophic link strength, connectance, and food chain lengths. There was evidence of an asymptotic relationship between energy flowing through a food chain and parasite diversity, although there were clear outliers. High parasite diversity was associated with host taxa which were highly connected within the food web. This suggests that energy flow through a taxon may favour parasite diversity, up to a maximal value. The evolutionary and energetic basis for that limitation is of key interest in understanding the basis for parasite diversity in natural food webs and thus their role in food web dynamics.

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Quantitative food web modeling unravels the importance of the microphytobenthos-meiofauna pathway for a high trophic transfer by meiofauna in soft-bottom intertidal food webs.
  • May 30, 2020
  • Ecological Modelling
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Quantitative food web modeling unravels the importance of the microphytobenthos-meiofauna pathway for a high trophic transfer by meiofauna in soft-bottom intertidal food webs.

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Evaluation of Daphnid Grazing on Microscopic Zoosporic Fungi by Using Comparative Threshold Cycle Quantitative PCR.
  • Apr 22, 2016
  • Applied and Environmental Microbiology
  • Michelle A Maier + 3 more

Lethal parasitism of large phytoplankton by chytrids (microscopic zoosporic fungi) may play an important role in organic matter and nutrient cycling in aquatic environments by shunting carbon away from hosts and into much smaller zoospores, which are more readily consumed by zooplankton. This pathway provides a mechanism to more efficiently retain carbon within food webs and reduce export losses. However, challenges in accurate identification and quantification of chytrids have prevented a robust assessment of the relative importance of parasitism for carbon and energy flows within aquatic systems. The use of molecular techniques has greatly advanced our ability to detect small, nondescript microorganisms in aquatic environments in recent years, including chytrids. We used quantitative PCR (qPCR) to quantify the consumption of zoospores by Daphnia in laboratory experiments using a culture-based comparative threshold cycle (CT) method. We successfully quantified the reduction of zoospores in water samples during Daphnia grazing and confirmed the presence of chytrid DNA inside the daphnid gut. We demonstrate that comparative CT qPCR is a robust and effective method to quantify zoospores and evaluate zoospore grazing by zooplankton and will aid in better understanding how chytrids contribute to organic matter cycling and trophic energy transfer within food webs. The study of aquatic fungi is often complicated by the fact that they possess complex life cycles that include a variety of morphological forms. Studies that rely on morphological characteristics to quantify the abundances of all stages of the fungal life cycle face the challenge of correctly identifying and enumerating the nondescript zoospores. These zoospores, however, provide an important trophic link between large colonial phytoplankton and zooplankton: that is, once the carbon is liberated from phytoplankton into the parasitic zoospores, the latter are consumed by zooplankton and carbon is retained in the aquatic food web rather than exported from the system. This study provides a tool to quantify zoospores and evaluate the consumption of zoospores by zooplankton in order to further our understanding of their role in food web dynamics.

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Pressure Loading of Piezo Composite Unimorphs
  • Jan 1, 2005
  • MRS Proceedings
  • Poorna Mane + 2 more

ABSTRACTOver the past decade synthetic jets have emerged as a promising means of active flow control. They have the ability to introduce small amounts of energy locally to achieve non-local changes in the flow field. These devices have the potential of saving millions of dollars by increasing the efficiency and simplifying fluid related systems. A synthetic jet actuator consists of a cavity with an oscillating diaphragm. As the diaphragm oscillates, jets are formed through an orifice in the cavity. This paper focuses on piezoelectric synthetic jets formed using two types of active diaphragms, Thunder® and Lipca. Thunder® is composed of three layers; two metal layers, with a PZT-5A layer in between, bonded with a polyimide adhesive. Lipca is a Light WeIght Piezo Composite Actuator, formed of a number of carbon fiber prepreg layers and an active PZT-5A layer. As these diaphragms oscillate, pressure differences within the cavity as well as average maximum jet velocities are measured. These parameters are measured under load and no-load conditions by controlling pressure at the back of the actuator or the passive cavity. Results show that the average maximum jet velocities measured at the exit of the active cavity, follow a similar trend to the active pressures for both devices. Active pressure and jet velocity increase with passive pressure to a maximum, and then decrease. Active pressure and the jet velocity peaked at the same passive cavity pressure of 18kPa for both diaphragms indicating that the same level of pre-stresses is present in both actuators even though Lipca produces approximately 10% higher velocities than Thunder®.

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  • Cite Count Icon 13
  • 10.1002/rra.1527
CAN WE IMPROVE MANAGEMENT PRACTICE OF FLOODPLAIN LAKES USING CLADOCERAN ZOOPLANKTON?
  • May 10, 2011
  • River Research and Applications
  • G R Kattel

ABSTRACTNatural resource managers across Australia intend to promote healthy floodplain lake ecosystems with rich diversity and composition of biota because such ecosystems provide economically valuable services to society. However, management practice of these floodplain lake ecosystems is impeded by confounding effects of anthropogenic impacts and natural climate variability in recent decades. Yet, there are a few potential biological markers available that profoundly respond to ecological effects of climate change and human disturbances. Cladoceran zooplankton plays an intermediary role in food web dynamics. They show distinct responses to changes in temperature and environmental perturbations, such as acidification, nutrient loading and salinization. The effects of temperature and land‐use changes on food web dynamics and water quality, in particular, are major concerns for shallow lowland large river floodplain lakes management in Australia. Information on zooplankton assemblages and diversity can help increase our understanding of ecological processes in a wide range of environmental exposures. The study of cladoceran fossils and their ephippia preserved in floodplain lake sediment has substantially furthered our understanding of species–environment relationships at different temporal and spatial scales and allowed us to develop powerful inference models for degraded floodplain lake ecosystems. This consequently defines a benchmark of a shift from a naturally intact ecosystem to an ecologically poor regime. In this paper, I have made an attempt to persuade wetland managers through application of contemporary and palaeocladoceran communities to improve management practice of floodplain lake ecosystems in Australia by providing a range of examples. Copyright © 2011 John Wiley & Sons, Ltd.

  • Research Article
  • Cite Count Icon 11
  • 10.1016/j.petrol.2020.107745
Simulation study on cutting transport in a horizontal well with hydraulic pulsed jet technology
  • Aug 15, 2020
  • Journal of Petroleum Science and Engineering
  • Shuyan Wang + 6 more

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  • Research Article
  • Cite Count Icon 25
  • 10.3389/fneur.2020.00213
Absent Arm Swing and Dual Tasking Decreases Trunk Postural Control and Dynamic Balance in People With Parkinson's Disease
  • Apr 17, 2020
  • Frontiers in Neurology
  • Tarique Siragy + 1 more

Introduction: Falling during walking is a common occurrence in people with Parkinson's disease and is closely associated with severe social and medical consequences. Recent evidence demonstrates that arm swing affects dynamic balance in healthy young adults; however, it remains unexamined what its effect is in people with Parkinson's disease, particularly when combined with a secondary dual task.Methods: Twenty people with Parkinson's disease (63.78 ± 8.97) walked with two arm swing conditions (absent and normal) with and without a secondary dual task. Data were collected on a split-belt treadmill CAREN Extended-System (Motek Medical, Amsterdam, NL). Average and standard deviations for trunk linear and angular velocity were calculated along with their instantaneous values (during foot strikes) in all three axes. Averages and coefficient of variations for step length, time, and width; margin of stability; and harmonic ratios were also calculated.Results: Compared with normal arm swing, absent arm swing reduced the least affected leg's average step length and increased its step length coefficient of variation while increasing step time coefficient of variation in the most affected leg. Further, absent arm swing reduced trunk anteroposterior instantaneous angular velocity (least affected leg) and reduced anteroposterior instantaneous linear velocity (bilaterally). For the vertical axis, absent arm swing increased the trunk's average angular velocity but reduced its instantaneous linear velocity and angular velocity standard deviation (least affected leg). Additionally, the margin of stability increased when the arms were absent (least affected leg). Alternatively, dual tasking reduced average step time (most affected leg) and increased the step width coefficient of variation (bilaterally). Additionally, dual tasking increased the mediolateral average angular velocity, instantaneous linear velocity standard deviation (bilaterally), and instantaneous angular velocity standard deviation (least affected leg). For the vertical axis, dual tasking increased average linear and angular velocity standard deviation as well as instantaneous angular velocity standard deviation (bilaterally).Conclusion: Findings suggest that participants attempted to control extraneous trunk movement (due to absent arm swing) through compensatory responses in both lower and upper extremities. However, participants appeared to predominately compensate on their least affected side. Contrastingly, modifying mediolateral foot placement appeared to be the main means of maintaining walking stability while dual tasking.

  • Research Article
  • Cite Count Icon 1
  • 10.1016/0009-2509(93)80353-r
Use of submerged jets in gas desorption from supersaturated solutions
  • Jan 1, 1993
  • Chemical Engineering Science
  • N Fidi + 3 more

Use of submerged jets in gas desorption from supersaturated solutions

  • Research Article
  • Cite Count Icon 2
  • 10.1111/jzo.13050
An experimental framework for quantifying the degree of intraguild predation in omnivorous food webs in the field
  • Jan 22, 2023
  • Journal of Zoology
  • G.‐C Hsu

Intraguild predation (IGP) is common in natural and human‐managed systems and plays a critical role in food web dynamics. Although studies have documented the occurrence of IGP across a wide range of predator taxa, quantitative understanding regarding the degree/intensity of IGP remains lacking. I propose an experimental framework combining controlled feeding trials and stable isotope analysis to quantify the degree of IGP in an omnivorous food web in the field consisting of a top predator, a mesopredator, and a shared prey. The degree of IGP is defined as the proportion (in number) of mesopredators consumed in the total diet (shared prey + mesopredator) of top predators. Feeding trials along with stable isotope analysis are used to construct a standard curve of the relationship between the diet composition of top predator and its nitrogen isotope signatures. The nitrogen isotope signatures of field‐collected top predator individuals are then analyzed and interpolated to the standard curve to estimate the degree of IGP in the field. The proposed framework leverages the strengths of different experimental approaches to study trophic interactions, providing a practical tool for quantifying IGP in a more accurate (controlled feeding trials and standard IGP curve) and realistic (stable isotope analysis of field samples) fashion. The current framework can be further extended to food webs involving more complex interactions (e.g., multiple shared prey) and complemented with other approaches (e.g., molecular gut content analysis) to capture a more complete picture of IGP dynamics in the field.

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