Abstract
Against the current inversion theory on the origin of chromatid bridges and fragments at meiotic anaphases, the present study introduces an alternative interpretation. It implies a possibility of recombinations of chromatids at the chiasma region as the consequence of normal crossing-over (Fig. E). At this moment the random reunion of chromatids is subjected to certain limitation by the cleavage of chromosome matrix. Thus the resultant configurations are three types, X, Y and O, as represented by Fig. C. The former two types relate to the formation of bridge and fragment at anaphases; the last implies the dissolution of the chiasma. If these three types are subjected to such a series of relationship as the present work has shown, this will be not only at variance with the inversion theory, but also strongly support my theory of crossing-over.
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