Abstract

1 Carnivore Ecology and Conservation Research Group, Tokyo University of Agriculture and Technology, Saiwaicho 3-5-8, Fuchu City, Tokyo 183-8509, Japan 2 Laboratory of Ecometry and Ecoinformatics, Graduate School of Natural Sciences, Nagoya City University, and Research Center for Biological Diversity, Nagoya City University, 1 Yamanohata, Mizuho-cho, Mizuho-ku, Nagoya, Aichi 467-8501, Japan 3 Laboratory of Applied Ecological Techniques, 3-2, Enokuma, Oita City, Oita 870-0889, Japan 4 The Nature Conservation Society of Japan, Mitoyo Bldg. 2F 1-16-10 Shinkawa, Chuo-ku, Tokyo 104-0033, Japan

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