Assessment of growth parameters in Pinus sylvestris L. stands

ABSTRACTLack of available workforce in motor-manual operations has led to increased interest in mechanized precommercial thinning (PCT) treatments. We evaluated 15 years of growth and yield development of a very dense (27,000 stems ha–1) balsam fir (Abies balsamea (L.) Mill) and black spruce (Picea mariana (Mill.) B.S.P.) stand after six PCT treatments: (1) motor-manual (MAN), (2) motor-manual with crop tree release (MAN-C), (3) mechanized (MEC), (4) mechanized with residual patches (MEC-P), (5) semi-mechanized (mechanized followed by motor-manual, MEC-S), and (6) unthinned control (CON). Compared to CON, both MAN and MEC-S increased height of balsam fir study trees by 53–56% and diameter at breast height (DBH) by 57–58%. MAN-C and MEC-S increased DBH of black spruce study trees by 15–31% over all other treatments. The proportion of merchantable basal area (BA) was higher in MAN (62%) and MEC-S (56%) than in CON (22%) and MEC (25%), without a reduction in total BA. PCT treatments that provided a uniform distribution of potential crop trees (MAN, MEC-S) performed better than MAN-C and MEC-P, while MEC yielded the poorest responses. Our findings highlight the need to develop new tailored mechanized systems for high wood production and low operational cost.

Corridor thinning can be an efficient method for extracting biomass from young stands, but its effects on subsequent productivity are uncertain. Therefore, its long-term effects were studied, using data obtained from two pre-commercial thinning (PCT) experiments (over 28 years) and nine thinning experiments (over 22 years). In the PCT experiments, thinning with total corridor areas of 0 (control), 57, 65, 73, 79 and 82% were compared to selective PCT leaving 1000 and 1400 stems/ha. In the thinning experiments corridor thinning (50% corridor area) and selective thinning from below (50% of basal area) were compared. No significant differences in diameter at breast height (DBH) were found between the corridor PCT and control treatments, but the control resulted in approximately 28–83% higher standing volumes/ha than the PCT treatments. Corridor PCT and thinning treatments resulted in higher stand stem density and lower mean DBH than selective treatments. No significant differences in volume growth, standing volume, mortality volume or height growth were detected between selective and corridor thinning or between most of the PCT treatments. The findings indicate that relatively large amounts of biomass could be extracted schematically, by early thinning instead of PCT, in young Scots pine stands without significant future yield losses.

Data from seven Scots pine (Pinus sylvestris L.) stands in central Sweden that had undergone pre-commercial thinning (PCT) were used to investigate the biological effect of PCT on the diameter at breast height (DBH) growth of the remaining trees. Two treatments were considered: a PCT treatment and a control (C) with no PCT. The DBH of the trees in each stand was measured on up to four occasions over 15 years. We examined mean DBH and DBH growth of the largest 300, 600, 900 and 1200 trees ha−1. Two methods of selecting the trees used to calculate the mean DBH values for each measurement occasion were considered: the actual mean DBH (Dma), which is based on the DBH distribution of the trees on the measurement occasion in question, and the genuine mean DBH (Dmg), which is based on the DBH distribution of the trees on the final measurement occasion of the study and therefore focuses on the same set of trees for all measurement occasions. There was no clear difference between the Dma- and Dmg-based DBH increments, but the Dmg values tended to be somewhat larger both for the C and PCT treatments. Over a 15-year period, the relative mean yearly increments (Dma/Dmg) for different tree size classes ranged from 0.91 to 0.98, with lower values for the C treatment and larger DBH size classes. We found that PCT promotes DBH growth: over a 15-year period, the increased growth amounted to about 2.0 mm year−1 compared to the trees in unthinned plots.

The objective of this paper was to investigate the impact of initial spacing and pre-commercial thinning (PCT) on tree growth, characteristics and stem quality in plantation-grown jack pine (Pinus banksiana Lamb.) after 58 years of growth. Four spacings including three initial spacings without PCT treatment, i.e., 1.52 × 1.52 m (5 × 5 ft), 2.13 × 2.13 m (7 × 7 ft) and 2.74 × 2.74 m (9 × 9 ft) and one initial spacing with PCT treatment (5 × 5 ft thinned to 7 × 7 ft at age 14), were chosen to analyze diameter at breast height (DBH) distribution, survival, and various tree and stem quality characteristics (e.g., DBH, total tree height, stem volume, crown size, branch, taper, sweep and wobbling). Tree mortality during the first 25 growth years was quite low for all four spacings, while after 25 years, survival for the four spacings rapidly declined. The decrease in survival for spacing 5 × 5 was the steepest, followed by spacings 7 × 7, 9 × 9 and finally the PCT-treated stand. DBH distributions for all four spacings were significantly different from each other, and five years after PCT, the 5 × 5 to 7 × 7 spacings had a similar DBH distribution to that of the constant 7 × 7 spacing. Mean DBHs differed significantly among the initial spacings. The 5 × 5 spacing produced significantly smaller trees than did the other three spacings through the years, and the 9 × 9 spacing produced significantly larger trees. PCT had positive effects on DBH growth, and yielded longer merchantable stems and better stem quality. PCT had little effect on crown development. Initial spacing significantly affected diameter growth, crown width and branch development, commercial stem length and volume, stem taper and stem deformation. Narrower spacing produced smaller but better quality stems. Spacing 7 × 7 had the worst stem quality in terms of overall sweep and wave-like transversal deformation (wobbling). A narrow initial spacing followed by a PCT treatment may provide a compromise solution for both tree growth and stem quality. Key words: Initial spacing, thinning, survival, tree growth, stem quality

The effects of different rates of precommercial thinning (PCT) intensity on the stand developmental parameters in natural Picea orientalis stands were investigated to determine the most suitable thinning rate. The present study was performed in Trabzon province located in the northeastern of Turkey. In PCT plots, four different levels of PCT intensity were applied including control (unthinned), light (1.3–1.5 m spacing), moderate (1.5–2.0 m spacing) and heavy (2.5–3.0 m spacing). The values of diameter and height were measured at the end of the 2010, 2012, 2014, 2016 and 2018 growing seasons. The results indicated statistically significant (p < 0.05) effects of PCT levels on the diameter, total stand volume and total basal area increments in all periods measured. On the other hand, significant differences were at 99% confidence level for 6 and 8-year height increments. Although the diameter increments increased with increasing levels of precommercial thinning, the height increments did not increase. Also, total basal area increments were higher than in the control in all precommercial thinning treatments, except for the heavy PCT. Considering the best results of measured growth parameters, the early results showed that the light and moderate PCT had the most favorable effect on stand development.

The effects of four precommercial thinning (PCT) treatments on an even-aged northern conifer stand in Maine were investigated by examining stand structure and composition 32 years after treatment. Replicated treatments applied in 1976 included: (1) control (no PCT), (2) row thinning (rowthin; 5-ft-wide row removal with 3-ft-wide residual strips), (3) row thinning with crop tree release (rowthin+CTR; 5-ft-wide row removal with crop tree release at 8-ft intervals in 3-ft-wide residual strips), and (4) crop tree release (CTR; release of selected crop trees at 8×8-ft intervals). PCT plots had more large trees and fewer small trees than the control in 2008. There were no other significant differences between the rowthin and control. The rowthin+CTR and CTR treatments had lower total and hardwood basal area (BA) and higher merchantable conifer BA than the control. CTR also resulted in more red spruce (Picea rubens [Sarg.]) and less balsam fir (Abies balsamea [L.]) than the other treatments. Although stand structures for rowthin+CTR and CTR were similar, the percentage of spruce in CTR was greater. Although the less-intensive rowthin+CTR treatment may provide many of the same benefits as CTR, the latter would be the preferred treatment if increasing the spruce component of a stand is an objective. Overall, early thinning treatments were found to have long-term effects on key stand attributes, even more than 30 years after treatment in areas with mixed species composition and moderate site potential.

Photosynthetic capacity, nutrient and water status following precommercial thinning in Anatolian black pine

Herbicide application and precommercial thinning (PCT) are common silvicultural treatments used across North America and Europe. Despite this widespread use, long-term growth and yield responses from controlled experiments that include both of these treatments are relatively rare. We used 40-year growth and yield responses of spruce–fir stands to various combinations of early herbicide and PCT in a long-term silvicultural experiment in central Maine, USA, to calibrate the Forest Vegetation Simulator (Northeast Variant). Using the calibrated model, we projected rotation-length outcomes for stand development, merchantable wood volumes, and stumpage-based financial returns. Projections indicated gains in total yield (17%–31%) from herbicide treatments at the end of the rotation (∼60 years postharvest) relative to untreated stands. Substantial increases in merchantable wood volume also were achieved with PCT. Twenty-four years after PCT, stand stumpage value averaged $907 USD·ha−1 higher than that for unthinned stands. Total yield and stumpage gains from PCT were projected to continue through the end of rotation. Highest stumpage values resulted from combined herbicide and PCT treatments, followed by PCT-only and then herbicide-only. At end of rotation, highest net present value (NPV) resulted from PCT, whether alone or in combination with herbicides. PCT and herbicide investments substantially improved the NPV relative to untreated stands when using discount rates of 2% and 4%, but not when using a 6% rate. Our results documented that good financial returns are possible over the long-term from early investments in herbicide and PCT treatments in Maine spruce–fir stands.

We investigated the effects of precommercial thinning (PCT) applied at different levels on the growth parameters of natural Anatolian black pine stand. The experimental site was located in the province of Isparta in southern Turkey. PCT was set up using a randomized complete block design with three replicates and three treatments (control, 2–2.5 m × 2–2.5 m and 3–3.5 m × 3–3.5 m spacing). The experimental plots were established in March 2014. At the end of the growing seasons for three years (2014–2016), growth parameters such as diameter, height and crown radius were measured. PCT was found to be significantly effective on diameter, height, individual tree basal area, and crown radius increment at the end of the third year (p < 0.05). While diameter, individual tree basal area and crown radius increment increased with increasing PCT intensity, height increment was higher in control than PCT. Although the PCT treatments applied with 3–3.5 m spacing affected individual tree development positively, we need long-term results on wood quality and yield. According to the short-term results, it is thought that it is better to lower the stand closeness gradually, so, it may be advisable to apply PCT by 2–2.5 m × 2–2.5 m spacing.

Early management of the regenerated seedlings shapes the future stand properties. To address these issues, pre-commercial thinning (PCT) and control treatments were applied to planted Norway spruce (Picea abies L. Karst) and naturally regenerated birch (Betula pendula Roth., Betula pubescens Ehrh.) stands in forest experiments in southern Sweden (lat. 56–57 N) containing 1.1–5.5 m tall saplings. The treatments were retention of 1000 or 2000 stems ha−1 of Norway spruce, with no birch or birch at 1000 stems ha−1. Treatments were replicated with and without annual removal of birch sprouts from stumps. The periodic annual increment (PAI) over five years was calculated for total stand volume and individual trees. The mean PAI of dominant trees was significantly higher both following all PCT treatments than controls, and following low rather than high-density PCT. Birch retention did not affect growth of the dominant trees but PAI was lower in plots with uncontrolled sprouting. The PAI of birch was significantly higher in low-density Norway spruce plots than in control plots and the high-density plots. The treatment response was significant even in stands with initial heights of only 1–2 m.

The net photosynthetic rate (A), stomatal conductance (gs), and transpiration rate (E), chlorophyll content, and midday xylem water potential (Ψmd) of Pinus nigra Arn. subsp. pallasiana (Lamb.) Holmboe trees were studied in a natural stand (Burdur-Aziziye, Turkey) for the first (2015), second (2016) and third (2017) years after precommercial thinning (PCT) to better understand the physiological and biochemical responses of trees in a semiarid region. In this study, three treatments (control, 2–2.5 × 2–2.5 m and 3–3.5 × 3–3.5 m spacing) were applied. Soil water content (SWC) and soil temperature (ST) was greater in both PCT treatments than in the control treatments. In general, the control treatment had a lower Ψmd compared with PCT treatments. A, gs, and E are generally higher than PCT treatments applied with 3–3.5 m spacing. The lowest gs and E were determined in the control trees. Ψmd and A were positively correlated with SWC, but negatively correlated with ST. The decrease in stress level, improvement in resources, and reduced competition for resources by fewer trees increased the A and gs in trees in the PCT treatments. Chlorophyll a, chlorophyll b, and total chlorophyll content are higher in the control treatments as compared to PCT treatments. SWC, ST, Ψmd, A, gs, E, and photosynthetic pigment showed seasonal changes. The two PCT treatments were generally similar in the ecophysiological changes in plots and a longer time is needed to decide for PCT treatments applied with 3–3.5 m spacing. According to the results obtained, it is recommended that PCT be applied with spacings of 2–2.5 × 2–2.5 m.

Models for quantifying the growth responses of black spruce (Picea mariana (Mill) BSP.) and jack pine (Pinus banksiana Lamb.) to precommercial thinning (PCT) treatments were developed. They accounted for the increased rate of stand development arising from PCT treatments through temporal adjustments to the species and site specific mean dominant height-age functions. Analytically, they utilized a relative height growth modifier consistent with observed density-dependent height repression effects. A phenotypic juvenile age-mature age correlation function was used to account for the intrinsic temporal decline in the magnitude of the PCT effect throughout the rotation. The resultant stand development patterns were in accord with theoretical and empirical expectations when the response models were integrated into algorithmic variants of structural stand density management models.

We examined the development of a 33-year experiment in spruce–fir stands that received nine herbicide treatments (applied aerially in 1977), with and without precommercial thinning (PCT) (applied in 1986). We tested two commonly held assumptions about the long-term effects of herbicide and PCT in mixedwood stands managed for softwoods: (i) herbicide release produces stands dominated by softwoods and (ii) PCT promotes both softwood dominance and merchantable volume. All herbicides were effective at releasing balsam fir ( Abies balsamea (L.) Mill.) and spruce ( Picea spp.) from overtopping shrubs and hardwoods that had naturally regenerated following clearcut harvesting in 1970. Glyphosate (Roundup), triclopyr amine (Garlon 3A), 2,4,5-T, and a mixture of 2,4,5-T and 2,4-D herbicide treatments were all effective at shifting long-term species composition to softwood dominance after 30 years, irrespective of the rates applied. Herbicide-treated stands that did not receive PCT were overstocked with softwoods for at least 24 years and produced less than half of the merchantable softwood volume by 40 years than herbicide-treated stands that received PCT. Intolerant hardwoods dominated untreated stands for all 33 years of the experiment. When unsprayed plots received PCT, however, they produced stands at 40 years that were compositionally and structurally indistinguishable from those that had received both herbicide and PCT treatments. Results from this experiment clearly demonstrated that early herbicide application, regardless of type and rate of herbicide, created softwood-dominated mixedwood stands over the long term and that PCT more than doubled merchantable softwood volume within 25 years of application.

Influence of stand thinning and repeated fertilization on plant community abundance and diversity in young lodgepole pine stands: 15-year results

The effects of precommercial thinning (PCT) on stem dimensions, form, volume, and branch attributes of red spruce [Picea rubens Sarg.] and balsam fir [Abies balsamea (L.) Mill.] crop trees were assessed 25 years after treatment in an even-aged northern conifer stand. Treatments were a uniform 2.4 x 2.4-m spacing and a control (no PCT). The PCT treatment significantly increased individual tree diameter at breast height (DBH), height growth, crown ratio, and crown width, while it reduced the tree height to DBH ratio. PCT also significantly increased stem taper and consequently, regional volume equations overpredicted observed stem volume by 2 to 15%, particularly for the spaced trees. PCT also increased the number and maximum size of branches on the lower bole. The sizes of knots on half of the sampled spruce crop trees in the spaced plots precluded them from being used as select structural lumber; there were no other effects on log grade. Our findings indicate that PCT can have a long-term influence on the structural attributes of individual trees, and that improved stem-volume prediction equations are needed in the Acadian region of North America.